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  • E-mailArild.Folkvord@uib.no
  • Phone+47 55 58 44 56+47 413 00 147
  • Visitor Address
    Thormøhlensgt. 53 A/B
    Room 
    2G19
  • Postal Address
    Postboks 7803
    5020 Bergen

The focus of my research is on growth and survival mechanisms in the early life of fishes. The research is experimentally oriented, and covers aspects of marine juvenile production and process oriented recruitment studies.

Size-selective mortality in cultured fish is often due to aggression and cannibalism. A large effort has been made to elucidate factors promoting cannibalism in captive cod (e.g. hunger level, size, size differences etc.). Size- and temperature dependent growth in juvenile flatfish and cod has also been studied with the aim of determining the growth potential under culture conditions. The growth process is very dynamic during the early life stages, and controlled laboratory experiments are carried out to calibrate methods used for growth characterization (e.g. RNA:DNA analysis, otolith microstructure analysis, fatty acid analysis).

The age of field caught fish larvae is often difficult to assess. Fortunately, growth increments are usually formed on a daily basis in the ear stones (otoliths) of fish larvae. It is thus possible to estimate the larval age in days in a similar manner as one determines the age of older fish by counting yearly zones in the otoliths and scales. The growth pattern in the otolith also reflects the previous growth history of the larva, and can be used to determine where, and under what conditions, the larva grew up. The otolith has therefore often been referred to as the "black box" in fish. An important part of the ageing research is related to the accuracy and precision of the age estimation. Knowledge in this field is of great importance in process studies and fisheries management.

I currently hold an adjunct position at Institute of Marine Research and am a member of the Hjort Center for Marine Ecosystem Dynamics.

Journal articles
  • Berg, Florian; Husebø, Åse; Godiksen, Jane Aanestad; Slotte, Aril; Folkvord, Arild. 2017. Spawning time of Atlantic herring (Clupea harengus) populations within a restricted area reflects their otolith growth at the larval stage. Fisheries Research. 194: 68-75. doi: 10.1016/j.fishres.2017.05.009
  • Blanco, Edurne; Reglero, Patricia; Ortega, Aurelio; de la Gándara, Fernando; Fiksen, Øyvind; Folkvord, Arild. 2017. The effects of light, darkness and intermittent feeding on the growth and survival of reared Atlantic bonito and Atlantic bluefin tuna larvae. Aquaculture. 479: 233-239. doi: 10.1016/j.aquaculture.2017.05.020
  • Feng, Chungang; Pettersson, Mats; Lamichhaney, Sangeet; Rubin, Carl-Johan; Rafati, Nima; Casini, Michele; Folkvord, Arild; Andersson, Leif. 2017. Moderate nucleotide diversity in the Atlantic herring is associated with a low mutation rate. eLIFE. 6:e23907: 1-14. doi: 10.7554/eLife.23907
  • Barrio, Alvaro Martinez; Lamichhaney, Sangeet; Fan, Guangyi; Rafati, Nima; Pettersson, Mats; Zhang, He; Dainat, Jacques; Ekman, Diana; Höppner, Marc P.; Jern, Patric; Martin, Marcel; Nystedt, Björn; Liu, Xin; Chen, Wenbin; Liang, Xinming; Shi, Chengcheng; Fu, Yuanyuan; Ma, Kailong; Zhan, Xiao; Feng, Chungang; Gustafson, Ulla; Rubin, Carl-Johan; Almén, Markus Sällman; Blass, Martina; Casini, Michele; Folkvord, Arild; Laikre, Linda; Ryman, Nils; Lee, Simon Ming-Yuen; Xu, Xun; Andersson, Leif. 2016. The genetic basis for ecological adaptation of the Atlantic herring revealed by genome sequencing. eLIFE. 5. doi: 10.7554/eLife.12081
  • Folkvord, Arild; Gundersen, Geir; Albretsen, Jon; Asplin, Lars; Kaartvedt, Stein; Giske, Jarl. 2016. Impact of hatch date on early life growth and survival of Mueller’s pearlside (Maurolicus muelleri) larvae and life-history consequences. Canadian Journal of Fisheries and Aquatic Sciences. 73: 163-176. doi: 10.1139/cjfas-2015-0040
  • Folkvord, Arild; Gundersen, Geir; Albretsen, Jon; Asplin, Lars; Kaartvedt, Stein; Giske, Jarl. 2016. Impact of hatch-date on early life growth and survival of Mueller's pearlside (Maurolicus Muelleri) larvae and life-history consequences. Canadian Journal of Fisheries and Aquatic Sciences. 73: 163-176. doi: 10.1139/cjfas-2015-0040
  • Folkvord, Arild; Vollset, Knut Wiik; Catalan, Ignacio. 2015. Differences in growth and survival between cod Gadus morhua and herring Clupea harengus early stages co-reared at variable prey concentrations. Journal of Fish Biology. 87: 1176-1190. doi: 10.1111/jfb.12783
  • Maneja, Rommel Hilot; Frommel, Andrea Y.; Browman, Howard; Geffen, Audrey J.; Folkvord, Arild; Piatkowski, Uwe; Durif, Caroline; Bjelland, Reidun Marie; Skiftesvik, Anne Berit; Clemmesen, Catriona M. 2015. The swimming kinematics and foraging behavior of larval Atlantic herring (Clupea harengus L.) are unaffected by elevated pCO2. Journal of Experimental Marine Biology and Ecology. 466: 42-48. doi: 10.1016/j.jembe.2015.02.008
  • Reglero, Patricia; Blanco, Edurne; Ortega, Aurelio; Fiksen, Øyvind; de la Gándara, Fernando; Seoka, Manabu; Viguri, F. Javier; Folkvord, Arild. 2015. Prey selectivity in piscivorous bluefin tuna larvae reared in the laboratory. Journal of Plankton Research. 37: 2-5. doi: 10.1093/plankt/fbu082
  • Folkvord, Arild; Jørgensen, Christian; Korsbrekke, Knut; Nash, Richard David Marriott; Nilsen, Trygve; Skjæraasen, Jon Egil. 2014. Trade-offs between growth and reproduction in wild Atlantic cod. Canadian Journal of Fisheries and Aquatic Sciences. 71: 1106-1112. doi: 10.1139/cjfas-2013-0600
  • Frommel, Andrea Y.; Maneja, Rommel; Lowe, David; Pascoe, Christine K.; Geffen, Audrey J.; Folkvord, Arild; Piatkowski, Uwe; Clemmesen, Catriona. 2014. Organ damage in Atlantic herring larvae as a result of ocean acidification. Ecological Applications. 24: 1131-1143. doi: 10.1890/13-0297.1
  • Hansen, Truls Wergeland; Folkvord, Arild; Grøtan, Espen; Sæle, Øystein. 2014. Corrigendum to "Genetic ontogeny of pancreatic enzymes in Labrus bergylta larvae and the effect of feed type on enzyme activities and gene expression" [Comp. Biochem. Physiol. B 164 (2013) 176-184]. Comparative Biochemistry and Physiology - Part B: Biochemistry & Molecular Biology. 170: 42. doi: 10.1016/j.cbpb.2014.01.005
  • Imsland, Albert; Ólafsson, Kristinn; Skírnisdóttir, Sigurlaug; Gunnarsson, Snorri; Oddgeirsson, Matthías; Vandamme, Sara; Helyar, Sarah J.; Skadal, Julie; Folkvord, Arild. 2014. Life history of turbot in Icelandic waters: Intra- and inter-population genetic diversity and otolith tracking of environmental temperatures. Fisheries Research. 155: 185-193. doi: 10.1016/j.fishres.2014.03.004
  • Jung, Kyung Mi; Folkvord, Arild; Kjesbu, Olav Sigurd; Sundby, Svein. 2014. Experimental parameterisation of principal physics in buoyancy variations of marine teleost eggs. PLoS ONE. 9. 10 pages. doi: 10.1371/journal.pone.0104089
  • Reglero, Patricia; Ortega, Aurelio; Blanco, Edurne; Fiksen, Øyvind; Viguri, F. Javier; de la Gándara, Fernando; Seoka, Manabu; Folkvord, Arild. 2014. Size-related differences in growth and survival in piscivorous fish larvae fed different prey types. Aquaculture. 433: 94-101. doi: 10.1016/j.aquaculture.2014.05.050
  • Hansen, Truls Wergeland; Folkvord, Arild; Grøtan, Espen; Sæle, Øystein. 2013. Genetic ontogeny of pancreatic enzymes in Labrus bergylta larvae and the effect of feed type on enzyme activities and gene expression. Comparative Biochemistry and Physiology - Part B: Biochemistry & Molecular Biology. 164: 176-184. doi: 10.1016/j.cbpb.2012.12.001
  • Karlson, Stine; Michalsen, Kathrine; Folkvord, Arild. 2013. Age determination of Atlantic halibut (Hippoglossus hippoglossus L.) along the coast of Norway: status, and improvements. ICES Journal of Marine Science. 70: 50-55. doi: 10.1093/icesjms/fss174
  • Maneja, Rommel Hilot; Frommel, Andrea Y.; Browman, Howard; Clemmesen, Catriona; Geffen, Audrey J.; Folkvord, Arild; Piatkowski, Uwe; Durif, Caroline; Bjelland, Reidun Marie; Skiftesvik, Anne Berit. 2013. The swimming kinematics of larval Atlantic cod, Gadus morhua L., are resilient to elevated seawater pCO2. Marine Biology. doi: 10.1007/s00227-012-2054-y
  • Maneja, Rommel; Frommel, Andrea Y.; Geffen, Audrey J.; Folkvord, Arild; Piatkowski, Uwe; Chang, Mei-Yu; Clemmesen, Catriona. 2013. Effects of ocean acidification on the calcification of otoliths of larval Atlantic cod Gadus morhua. Marine Ecology Progress Series. 477: 251-258. doi: 10.3354/meps10146
  • Ritchie, Kenneth; Folkvord, Arild; Hufthammer, Anne Karin. 2013. Oxygen isotope ratios in cod otoliths used to reveal seasonality of fishing at Late Mesolithic sites in Denmark. Archaeofauna. 22: 95-104.
  • Vollset, Knut Wiik; Catalan, IA; Fiksen, Øyvind; Folkvord, Arild. 2013. Effect of food deprivation on distribution of larval and early juvenile cod in experimental vertical temperature and light gradients. Marine Ecology Progress Series. 475: 191-201. doi: 10.3354/meps10129
  • Ferreira, Ana Sofia de Araújo; Stenevik, Erling Kåre; Vollset, Knut Wiik; Korneliussen, Rolf; Folkvord, Arild. 2012. Vertical migration of Norwegian spring-spawning herring larvae in relation to predator and prey distribution. Marine Biology Research. 8: 605-614. doi: 10.1080/17451000.2011.653370
  • Frommel, Andrea Y.; Maneja, Rommel; Lowe, David; Malzahn, Arne M.; Geffen, Audrey J.; Folkvord, Arild; Piatkowski, Uwe; Reusch, Thorsten B.H.; Clemmensen, Catriona. 2012. Severe tissue damage in Atlantic cod larvae under increasing ocean acidification. Nature Climate Change. 2: 42-46. doi: 10.1038/nclimate1324
  • Jung, Kyung Mi; Folkvord, Arild; Kjesbu, Olav Sigurd; Agnalt, Ann-Lisbeth; Thorsen, Anders; Sundby, Svein. 2012. Egg buoyancy variability in local populations of Atlantic cod (Gadus morhua). Marine Biology. 159: 1969-1980. doi: 10.1007/s00227-012-1984-8
  • Koedijk, Roland; Imsland, Albert; Folkvord, Arild; Stefansson, Sigurd Olav; Jonassen, Thor Magne; Foss, Atle. 2012. Larval rearing environment influences the physiological adaptation in juvenile Atlantic cod, Gadus morhua. Aquaculture International. 20: 467-479. doi: 10.1007/s10499-011-9478-0
  • Meyer, Stefan; Caldarone, Elaine M.; Chicharo, Maria Alexandra; Clemmesen, Catriona; Faria, Ana M.; Faulk, Cynthia K.; Folkvord, Arild; Holt, G. Joan; Høie, Hans; Kanstinger, Philipp; Malzahn, Arne; Moran, Damian; Petereit, Christoph; Støttrup, Josianne G.; Peck, Myron A. 2012. On the edge of death: Rates of decline and lower thresholds of biochemical condition in food-deprived fish larvae and juveniles. Journal of Marine Systems. 93: 11-24. doi: 10.1016/j.jmarsys.2011.09.010
  • Catalan, Ignacio; Vollset, Knut Wiik; Morales-Nin, Beatriz; Folkvord, Arild. 2011. The effect of temperature gradients and stomach fullness on the vertical distribution of larval herring in experimental columns. Journal of Experimental Marine Biology and Ecology. 404: 26-32. doi: 10.1016/j.jembe.2011.05.005
  • Geffen, Audrey J.; Høie, Hans; Folkvord, Arild; Hufthammer, Anne Karin; Andersson, Carin; Ninnemann, Ulysses S; Pedersen, Rolf B.; Nedreaas, Kjell. 2011. High-latitude climate variability and its effect on fisheries resources as revealed by fossil cod otoliths. ICES Journal of Marine Science. 68: 1081-1089. doi: 10.1093/icesjms/fsr017
  • Imsland, Albert; Koedijk, Roland; Stefansson, Sigurd Olav; Foss, Atle; Hjorleifsdottir, Sigrid; Hreggvidsson, Gudmundur O.; Otterlei, Erling; Folkvord, Arild. 2011. A retrospective approach to fractionize variation in body mass of Atlantic cod Gadus morhua. Journal of Fish Biology. 78: 251-264. doi: 10.1111/j.1095-8649.2010.02860.x
  • Vollset, Knut Wiik; Folkvord, Arild; Browman, Howard. 2011. Foraging behaviour of larval cod (Gadus morhua) at low light intensities. Marine Biology. 158: 1125-1133. doi: 10.1007/s00227-011-1635-5
  • Catalan, Ignacio; Folkvord, Arild; Palomera, Isabel; Quilez-Badia, Gemma; Kallianoti, Fotini; Tselepides, Anastasios; Kallianotis, Argyris. 2010. Growth and feeding patterns of European anchovy (Engraulis encrasicolus) early life stages in the Aegean Sea (NE Mediterranean). Estuarine, Coastal and Shelf Science. 86: 299-312. doi: 10.1016/j.ecss.2009.11.033
  • Folkvord, Arild; Koedijk, Roland; Lokøy, Vibeke Saure; Imsland, Albert. 2010. Timing and selectivity of mortality in reared Atlantic cod revealed by otolith analysis. Environmental Biology of Fishes. 89: 513-519. doi: 10.1007/s10641-010-9699-5
  • Hufthammer, Anne Karin; Høie, Hans; Folkvord, Arild; Geffen, Audrey J.; Andersson, Carin; Ninnemann, Ulysses S. 2010. Seasonality of human site occupation based on stable oxygen isotope ratios of cod otoliths. Journal of Archaeological Science. 37: 78-83. doi: 10.1016/j.jas.2009.09.001
  • Koedijk, Roland; Folkvord, Arild; Foss, Atle; Pittman, Karin; Stefansson, Sigurd Olav; Handeland, Sigurd O; Imsland, Albert. 2010. The influence of first-feeding diet on the Atlantic cod Gadus morhua phenotype: survival, development and long-term consequences for growth. Journal of Fish Biology. 77: 1-19. doi: 10.1111/j.1095-8649.2010.02652.x
  • Koedijk, Roland; Le François, Nathalie R.; Blier, Pierre U.; Foss, Atle; Folkvord, Arild; Ditlecadet, Delphine; Lamarre, Simon G.; Stefansson, Sigurd Olav; Imsland, Albert. 2010. Ontogenetic effects of diet during early development on growth performance, myosin mRNA expression and metabolic enzyme activity in Atlantic cod juveniles reared at different salinities. Comparative Biochemistry and Physiology A. 156: 102-109. doi: 10.1016/j.cbpa.2010.01.007
  • Meier, Sonnich; Morton, Craig H.; Nyhammer, Gunnar; Grøsvik, Bjørn Einar; Makhotin, Valeri; Geffen, Audrey J.; Boitsov, Stephan; Kvestad, Karen Anita; Kjersem, Anneli Bohne; Goksøyr, Anders; Folkvord, Arild; Klungsøyr, Jarle; Svardal, Asbjørn M. 2010. Development of Atlantic cod (Gadus morhua) exposed to produced water during early life stages: Effects on embryos, larvae, and juvenile fish. Marine Environmental Research. 70: 383-394. doi: 10.1016/j.marenvres.2010.08.002
  • Seljeset, Oddbjørn; Vollset, Knut Wiik; Folkvord, Arild; Geffen, Audrey J. 2010. The role of prey concentration and size range in the growth and survival of larval cod. Marine Biology Research. 6: 251-262. doi: 10.1080/17451000903150355
  • Busch, Kjersti Eline Tønnessen; Folkvord, Arild; Otterå, Håkon; Hutchinson, William F.; Svåsand, Terje. 2009. Effects of female spawning experience and larval size on feeding and growth of cod larvae Gadus morhua L. reared in mesocosms. Marine Biology Research. 5: 286-296. doi: 10.1080/17451000802441319
  • Folkvord, Arild; Fiksen, Øyvind; Høie, Hans; Johannessen, Arne; Otterlei, Erling; Vollset, Knut Wiik. 2009. What can size distributions within cohorts tell us about ecological processes in fish larvae? Scientia Marina. 73: 119-130. doi: 10.3989/scimar.2009.73s1119
  • Folkvord, Arild; Høie, Hans; Johannessen, Arne; Solbakken, Turid. 2009. Effects of prey concentration, light regime, and parental origin on growth and survival of herring larvae under controlled experimental conditions. ICES Journal of Marine Science. 66: 1702-1709. doi: 10.1093/icesjms/fsp072
  • Husebø, Åse; Stenevik, Erling Kåre; Slotte, Aril; Fossum, Petter; Salthaug, Are; Vikebø, Frode Bendiksen; Aanes, Sondre; Folkvord, Arild. 2009. Effects of hatching time on year-class strength in Norwegian spring-spawning herring (Clupea harengus). ICES Journal of Marine Science. 66: 1710-1717. doi: 10.1093/icesjms/fsp150
  • Vollset, Knut Wiik; Fiksen, Øyvind; Folkvord, Arild. 2009. Vertical distribution of larval cod (Gadus morhua) in experimental temperature gradients. Journal of Experimental Marine Biology and Ecology. 379: 16-22. doi: 10.1016/j.jembe.2009.08.013
  • Vollset, Knut Wiik; Seljeset, Oddbjørn; Fiksen, Øyvind; Folkvord, Arild. 2009. A common garden experiment of larval Northeast Arctic and Norwegian coastal cod cohorts in replicated mesocosms. Deep-sea research. Part II, Topical studies in oceanography. 56: 1984-1991. doi: 10.1016/j.dsr2.2008.11.009
  • Høie, Hans; Folkvord, Arild; Mosegaard, Henrik; Li, Lingbo; Clausen, Lise A. Worsøe; Norberg, Birgitta; Geffen, Audrey. 2008. Restricted fish feeding reduces cod otolith opacity. Journal of Applied Ichthyology. 24: 138-143. doi: 10.1111/j.1439-0426.2007.01014.x
  • Li, Lingbo; Høie, Hans; Geffen, Audrey J.; Heegaard, Einar; Skadal, Julie; Folkvord, Arild. 2008. Back-calculation of previous fish size using individually tagged and marked Atlantic cod (Gadus morhua). Canadian Journal of Fisheries and Aquatic Sciences. 65: 2496-2508. doi: 10.1139/F08-157
  • Urtizberea, Agurtzane; Fiksen, Øyvind; Folkvord, Arild; Irigoien, Xabier. 2008. Modelling growth of larval anchovies including diel feeding patterns, temperature and body size. Journal of Plankton Research. 30: 1369-1383. doi: 10.1093/plankt/fbn090
  • Bang, Anders; Grønkjær, Peter; Folkvord, Arild. 2007. Possible fitness costs of high and low standard metabolic rates in larval herring Clupea harengus, as determined by otolith microstructure. Marine Ecology Progress Series. 331: 233-242.
  • Folkvord, Arild. 2007. Erratum: Comparison of size-at-age of larval Atlantic cod (Gadus morhua) from different populations based on size- and temperature-dependent grwoth models. Can. J. Fish. Aquat. Sci. 64: 583-585. Canadian Journal of Fisheries and Aquatic Sciences. 64: 583-585.
  • González-Quirós, Rafael; Munuera, Itziar; Folkvord, Arild. 2007. Cell cycle analysis of brain cells as a growth index in larval cod at different feeding conditions and temperatures. Scientia Marina. 71: 485-497.
  • Imsland, Albert; Foss, A; Koedijk, Roland; Folkvord, Arild; Stefansson, Sigurd; Jonassen, TM. 2007. Persistent growth effects of temperature and photoperiod in Atlantic cod Gadus morhua. Journal of Fish Biology. 71: 1371-1382. doi: 10.1111/j.1095-8649.2007.01600.x
  • Imsland, Albert; Foss, Atle; Alvseike, Thomas; Folkvord, Arild; Stefansson, Sigurd; Jonassen, Thor Magne. 2007. Interaction between temperature and photoperiod on growth and feeding of Atlantic cod (Gadus morhua): possible secondary effects. Canadian Journal of Fisheries and Aquatic Sciences. 64: 239-248. doi: 10.1139/F07-006
  • Kristiansen, Trond; Fiksen, Øyvind; Folkvord, Arild. 2007. Modelling feeding, growth, and habitat selection in larval Atlantic cod (Gadus morhua): observations and model predictions in a macrocosm environment. Canadian Journal of Fisheries and Aquatic Sciences. 64: 136-151. Published 2007-01-30. doi: 10.1139/F06-176
  • Tian, Tian; Fiksen, Øyvind; Folkvord, Arild. 2007. Estimating larval fish growth under size-dependent mortality: a numerical analysis of bias. Canadian Journal of Fisheries and Aquatic Sciences. 64: 554-562. doi: 10.1139/F07-031
  • Caldarone, Elaine M.; Clemmesen, Catriona M.; Berdalet, Elisa; Miller, Thomas J.; Folkvord, Arild; Holt, G. Joan; Olivar, M. Pilar; Suthers, Iain M. 2006. Intercalibration of four spectrofluorometric protocols for measuring RNA/DNA ratios in larval and juvenile fish. Limnology and Oceanography : Methods. 4: 153-163.
  • Høie, Hans; Folkvord, Arild. 2006. Estimating the timing of growth rings in Atlantic cod otoliths using stable oxygen isotopes. Journal of Fish Biology. 68: 826-837. doi: Se sammendrag
  • Imsland, Albert; Foss, A.; Folkvord, Arild; Stefansson, Sigurd; Jonassen, T.M. 2006. The interrelation between temperature regimes and fish size in juvenilie Atlantic cod (Gadus morhua). effects on growth and feed conversion efficiency. Fish Physiology & Biochemistry. 31: 347-361. doi: 10.1007/s10695-005-4244-8
  • Imsland, Albert; Foss, Atle; Koedijk, Roland; Folkvord, Arild; Stefansson, Sigurd; Jonassen, Thor M. 2006. Short- and long-term differences in growth, feed conversion efficiency and deformities in juvenile Atlantic cod (Gadus morhua) startfed on rotifers or zooplankton. Aquaculture Research. 37: 1015-1027.
  • Meisfjord, Jørgen Rajan; Midtøy, Frank; Folkvord, Arild. 2006. Validation of daily increment deposition in otoliths of juvenile Limnothrissa miodon (Clupeidae). Journal of Fish Biology. 69: 1845-1848.
  • Meisfjord, Jørgen Rajan; Midtøy, Frank; Folkvord, Arild. 2006. Validation of daily increment deposition in otoliths of juvenile Limnothrissa miodon (Clupeidae). Journal of Fish Biology. 69: 1845-1848. doi: 10.111/j.1095-8649.2006.01204
  • Simonsen, Claus S.; Munk, Peter; Folkvord, Arild; Pedersen, Søren Anker. 2006. Feeding ecology of Greenland halibut and sandeel larvae off West Greenland. Marine Biology. 149: 937-952. doi: 10.1007/s00227-005-0172-5
  • Folkvord, Arild. 2005. Comparison of size-at-age of larval Atlantic cod (Gadus morhua) from different populations based on size- and temperature-dependent growth models. Canadian Journal of Fisheries and Aquatic Sciences. 62: 1037-1052.
  • Imsland, Albert; Foss, Atle; Folkvord, Arild; Stefansson, Sigurd. 2005. Genotypic response to photoperiod treatment in Atlantic cod (Gadus morhua). Aquaculture. 250: 525-532.
  • Imsland, Albert; Foss, Atle; Folkvord, Arild; Stefansson, Sigurd; Jonassen, Thor Magne. 2005. The interrelation between temperature regimes and fish size in juvenile Atlantic cod (Gadus morhua): effects on growth and feed conversion efficiency. Fish Physiology & Biochemistry. 31: 347-361. doi: 10.10077s10695-005-4244-8
  • Belchier, M.; Clemmensen, C.; Cortés, D.; Doan, A.; Folkvord, Arild; García, A.; Geffen, Audrey; Høie, Hans; Johannessen, Arne; Moksness, E.; Pontual, E. de; Rámirez, T.; Sveinsbø, Bjørn. 2004. ICES. Recruitment studies: Manual of precision and accurac of tools. ICES Techniques in Marine Environmental Science. 33. 35 pages.
  • Folkvord, Arild; Johannessen, Arne; Moksness, Erlend. 2004. Temperature-dependent otolith growth in Norwegian spring-spawning herring (Clupea harengus L.) larvae. Sarsia. 89: 297-310.
  • Fox, Clive J; Folkvord, Arild; Geffen, Audrey. 2004. Otolith micro-increment formation in herring Clupea harengus larvae in relation to growth rate. Marine Ecology Progress Series. 264: 83-94.
  • Høie, Hans; Andersson, Carin; Folkvord, Arild; Karlsen, Ørjan. 2004. Precision and accuracy of stable isotope signals in otoliths of pen-reared cod (Gadus morhua) when sampled with a high-resolution micromill. Marine Biology. 144: 1039-1049. doi: 10.1007/s00227-003-1275-5
  • Høie, Hans; Otterlei, Erling; Folkvord, Arild. 2004. Temperature-dependent fractionation of stable oxygen isotopes in otoliths of juvenile cod (Gadus morhua L.). ICES Journal of Marine Science. 61: 243-251. doi: 10.1016/j.icesjms.2003.11.006
  • Imsland, Albert; Foss, Atle; Nævdal, Gunnar; Johansen, T.; Folkvord, Arild; Stefansson, Sigurd; Jonassen, Thor M. 2004. Variations in growth in haemoglobin genotypes of Atlantic cod. Fish Physiology & Biochemistry. 30: 47-55.
  • Skajaa, Katrine; Fernø, Anders; Folkvord, Arild. 2004. Ontogenetic- and condition-related effects of starvation on responsiveness in herring larvae (Clupea harengus L.) during repeated attacks by a model predator. Journal of Experimental Marine Biology and Ecology. 312: 253-269.
  • Otterlei, Erling; Folkvord, Arild; Nyhammer, Gunnar. 2002. Temperature dependent otolith growth of larval and early juvenile cod cod (Gadus morhua). ICES Journal of Marine Science. 59: 851-860.
  • Tytler, Peter J.; Fox, Clive J.; Folkvord, Arild. 2002. Glycoconjugates in the otolithic membranes of the larvae of herring: a framework for encoding the life history recorder in fish? Journal of Fish Biology. 61: 39-49.
  • Butler, John L.; Folkvord, Arild. 2000. Editorial. Section 2: Estimation of fish age and growth. Fisheries Research. 46: 51-52.
  • Folkvord, Arild; Blom, Geir; Johannessen, Arne; Moksness, Erlend. 2000. Growth-dependent age estimation in herring (Clupea harengus L.) larvae. Fisheries Research. 46: 91-103.
  • Johannessen, Arne; Blom, G.; Folkvord, Arild. 2000. Differences in otolith and somatic growth between spring and autumn spawned herring (Clupea harengus L.) larvae. Sarsia. 85: 461-466.
  • Fiksen, Øyvind; Folkvord, Arild. 1999. Modelling growth and ingestion processes in herring Clupea harengus larvae. Marine Ecology Progress Series. 184: 273-289.
  • Høie, Hans Ingolf; Folkvord, Arild; Johannessen, Arne. 1999. Maternal, paternal and temperature effects on otolith size of young herring (Clupea harengus L.) larvae. Journal of Experimental Marine Biology and Ecology. 234: 167-184.
  • Høie, Hans Ingolf; Folkvord, Arild; Johannessen, Arne. 1999. The influence of different parental combinations and incubation temperature on the RNA and DNA content of herring larvae at hatching: a pilot study. Journal of Fish Biology. 55: 110-118.
  • Kristogu-Baduge, Suneetha; Folkvord, Arild; Johannessen, Arne. 1999. Responsiveness of selected condition measures of herring, Clupea harengus, larvae to starvation in relation to ontogeny and temperature. Environmental Biology of Fishes. 54: 191-204.
  • Otterlei, Erling; Nyhammer, Gunnar; Folkvord, Arild; Stefansson, Sigurd. 1999. Temperature- and size-dependent growth of larval and early juvenile Atlantic cod (Gadus morhua): a comparative study of Norwegian coastal cod and northeast Arctic cod. Canadian Journal of Fisheries and Aquatic Sciences. 56: 2099-2111.
  • Imsland, A.K.; Nilsen, Trygve; Folkvord, Arild. 1998. Stochastic simulation of size variation in turbot: possible causes analysed with an individual-based model. Journal of Fish Biology. 53: 237-258.
  • Sunde, Leif M.; Imsland, Albert; Folkvord, Arild; Stefansson, Sigurd. 1998. Effects of size grading on growth and survival of juvenile turbot at two temperatures. Aquaculture International. 6: 19-32.
  • Bergstad, Odd Aksel; Folkvord, Arild. 1997. Dispersal of tagged juvenile turbot Scophthalmus maximus on the Norwegian Skagerrak coast. Fisheries Research. 29: 211-215.
  • Blom, Geir; Folkvord, Arild. 1997. A snapshot of cannibalism in 0-group Atlantic cod (Gadus morhua) in a marine pond. Journal of Applied Ichthyology. 13: 177-181.
  • Blom, Geir; Folkvord, Arild; Johannessen, Arne; Fossum, P. 1997. Interannual variations in condition indices of larval Norwegian spring-spawning herring. Journal of Fish Biology. 51 (Suppl A): 370-384.
  • Folkvord, Arild; Rukan, K.; Johannessen, Arne; Moksness, E. 1997. Early life history of herring (Clupea harengus L.) larvae in contrasting environments determined by otolith microstructure analysis. Journal of Fish Biology. 126: 250-263.
  • Imsland, A.K.; Folkvord, Arild; Grung, G.L.; Stefansson, S.O.; Taranger, G.L. 1997. Sexual dimorphism in growth and maturation of turbot, Scophthalmus maximus (Rafinesque, 1810). Aquaculture Research. 28: 101-114.
  • Imsland, Albert; Folkvord, Arild; Jónsdóttir, Ólöf D.B.; Stefansson, Sigurd. 1997. Effects of exposure to extended photoperiods during the first winter on long-term growth and age at first maturity in turbot (Scophthalmus macimus). Aquaculture. 159: 125-141.
  • Folkvord, Arild; Haaland, Inger Margrethe; Grahl-Nielsen, Otto; Johannesen, Arne. 1996. Fatty acid composition of herring (Clupea harengus L.) larvae off western Norway. Sarsia. 80: 299-305.
  • Folkvord, Arild; Ystanes, L.; Johannessen, A.; Moksness, E. 1996. RNA : DNA ratios and growth of herring (Clupea harengus) larvae reared in mesocosms. Marine Biology. 126: 591-602.
  • Imsland, A.K.; Sunde, L.M.; Folkvord, Arild; Stefansson, S.O. 1996. The interaction of temperature and fish size on growth of juvenile turbot. Journal of Fish Biology. 49: 926-940.
  • Stenevik, E. K.; Fossum, P.; Johannessen, A.; Folkvord, Arild. 1996. Identification of Norwegian spring spawning herring (Clupea harengus L .) larvae from spawning grounds off western Norway applying otolith microstructure analysis. Sarsia. 80: 285-292.
  • Folkvord, Arild; Moksness, Erlend. 1995. RNA/DNA ratios and growth of herring larvae. Marine Ecology Progress Series. 212: 311-312.
  • Hallaråker, H.; Folkvord, Arild; Stefansson, Sigurd. 1995. Growth of juvenile halibut (Hippoglossus hippoglossus L.) related to temperature, day length and feeding regime. Netherlands Journal of Sea Research. 34: 139-147.
  • Imsland, A. K.; Folkvord, Arild; Stefansson, Sigurd. 1995. Growth, oxygen consumption and activity of juvenile turbot (Scophthalamus maximus) reared under different temperatures and photoperiods. Netherlands Journal of Sea Research. 34: 149-159.
  • Folkvord, Arild; Blom, G.; Dragesund, O.; Johannessen, A.; Nakken, O.; Nævdal, Gunnar. 1994. A conceptual framework for enhancing and studying recruitment of marine fish stocks. Aquaculture and Fisheries Management. 25: 245-258.
  • Folkvord, Arild; Øiestad, V.; Kvenseth, P. G. 1994. Growth patterns of three cohorts of Atlantic cod larvae (Gadus morhua L.) studied in a macrocosm. ICES Journal of Marine Science. 51: 325-336.
  • Otterlei, E.; Folkvord, Arild; Møller, D. 1994. Effects of temperature and density on growth, survival, and cannibalism of juvenile cod (Gadus morhua L.). (Abstract). ICES Marine Science Symposia. 198: 632.
  • Folkvord, Arild. 1993. Prey recognition in stomachs of cannibalistic juvenile cod (Gadus morhua L.). Sarsia. 78: 4.
  • Folkvord, Arild; Otterå, Håkon. 1993. Effects of initial size distribution, day length, and feeding frequency on growth, survival, and cannibalism in juvenile Atlantic cod (Gadus morhua L.). Aquaculture. 114: 243-260.
  • Otterå, Håkon; Folkvord, Arild. 1993. Allometric growth in juvenile cod (Gadus morhua) and possible effects on cannibalism. Journal of Fish Biology. 43: 643-645.
  • Nævdal, Gunnar; Folkvord, Arild; Otterlei, E.; Thorkildsen, Solveig. 1992. Growth rate related to genotype of o-group cod at three environmental temperatures. Sarsia. 77: 71-73.
  • Booman, Clelia; Folkvord, Arild; Hunter, J. R. 1991. Responsivness of starved northern anchory larvae to predatory attacks by adult anchory. Fishery Bulletin U.S.. 89: 707-711.
  • Folkvord, Arild. 1991. Growth, survival and cannibalism of cod juveniles (Gadus morhua L.): effects of feed type, starvation and fish size. Aquaculture. 97: 41-59.
Reports and theses
  • Sundby, Svein; Fossum, Petter; Sandvik, Anne Dagrun; Vikebø, Frode Bendiksen; Aglen, Asgeir; Buhl-Mortensen, Lene; Folkvord, Arild; Bakkeplass, Kjell Gunnar; Buhl-Mortensen, Pål; Johannessen, Magnus Einar; Jørgensen, Martin S.; Kristiansen, Trond; Landa, Camilla S.; Myksvoll, Mari Skuggedal; Nash, Richard David Marriott. 2013. KunnskapsInnhenting Barentshavet-Lofoten-Vesterålen (KILO). Status og videreutvikling av kunnskapsgrunnlaget for mengde, fordeling og transport av kommersielle fiskearter fra gytende fisk til egg, larver, yngel og ungfisk i nordnorske sokkelområder med vekt på Lofoten og Vesterålen. Fisken og Havet, Særnummer. 3. Havforskningsinstituttet, Bergen. 186 pages.
  • Fiksen, Øyvind; Otterlei, Erling; Folkvord, Arild. 2000. Experiments and models as reciprocal tools to understand environmental links in recruitment dynamics. ICES CM Documents. R:02. International Council for the Exploration of the Sea, København. 8 pages.
  • Høie, Hans Ingolf; Folkvord, Arild; Johannessen, Arne. 2000. A multivariate analysis of condition of herring larvae from different environmental conditions. ICES CM Documents. R:04. ICES, København. 10 pages.
  • Folkvord, Arild; Otterlei, Erling; Svardal, Asbjørn. 1999. Alkylerte fenolers hormonelle innvirkning på torsk - generasjonseffekter. Vekst, overlevelse og kjønnsdifferensiering. IFM rapport. 15. Institutt for fiskeri- og marinbiologi, Univ. i Bergen, Bergen. 11 pages.
  • Folkvord, Arild; Blom, Geir; Helvik, Jon Vidar; Holmqvist, Bo I.; Johannessen, Arne; Otterlei, Erling; Stefansson, Sigurd; Brattegard, Torleiv. 1998. Teori- og eksperimentbasert studie av rekruttering hos marin fisk. Sluttrapport for del I av Strategisk Program. IFM rapport. 7. Inst. for fiskeri- og marinbiologi, Univ. i Bergen. 46 pages.
  • Folkvord, Arild; Johannessen, Arne; Brattegard, Torleiv; Nylund, Are. 1998. Otolitt mikrostruktur hos sild, presisjon, nøyaktighet og anvendelse. IFM rapport. 1. Inst. for fiskeri- og marinbiologi, Univ. i Bergen. 19 pages.
  • Folkvord, Arild. 1993. Experimental studies of growth and mortality of larval and juvenile cod (Gadus morhua L.) with special emphasis on cannibalism. Institutt for fiskeri- og marinbiologi. 153 pages.
  • Johannessen, Arne; Blom, Geir; Folkvord, Arild. 1993. Toktrapport. IFM rapport. 42. [Mangler utgivernavn]. 20 pages.
  • Folkvord, Arild; Johannessen, A. 1992. Toktrapport M/S Håkon Mosby, 4-13 april 1992. Universitetet i Bergen, inst. for fiskeri- og marinbiologi. 14 pages.
Book sections
  • Skajaa, Katrine; Fernö, Anders; Folkvord, Arild. 2003. Swimming, feeding and predator avoidance in cod larvae (Gadus morhua L.): trade offs between hunger and predation risk. Artikkel, pages 105-121. In:
    • Browman, Howard I.; Skiftesvik, Anne Berit. 2003. The Big Fish Bang. Proceedings of the 26th Annual Larval Fish Conference. 2003. ISBN: 82-7461-059-8.
  • Folkvord, Arild. 1997. Chapter nine. Ontogeny of cannibalism in larval and juvenile fishes with special emphasis on Atlantic cod. 251-278. In:
    • Chambers, R. Christopher; Trippel, Edward A. 1997. Early Life History and Recruitment in Fish Populations. ISBN: 0-412-64190-9.
  • Johannessen, A.; Blom, G.; Folkvord, Arild; Svendsen, Harald. 1995. The effect of local wind on the distribution of early Norwegian spring spawning herring (Clupea harengus L.) larvae. 365-384. In:
    • Skjoldal, H.R.; Hopkins, C.; Erikstad, K.E.; Leinaas, H.P. 1995. Ecology of Fjords and Coastal Waters.
  • Moksness, E.; Rukan, K.; Ystanes, L.; Folkvord, Arild; Johannessen, A. 1995. Comparison of somatic and otolith growth in North Sea herring (Clupea harengus L.) larvae; evaluation of growth dynamics in mesocosms. 119-134. In:
    • Secor, D.H.; Dean, J.M.; Campana, S. 1995. Recent Developments in Fish Otolith Research. The Belle W. Baruch Library in Marine Science.
  • Folkvord, Arild; Otterå, H. 1991. Effect of size distribution, feeding and light regime on growth and survival of cod juveniles in tanks. 299-300. In:
    • Lavens, P.; Sorgeloos, P.; Jaspers, E.; Ollevier, F. 1991. EAS Spec. Publ.

More information in national current research information system (CRIStin)

 

Link to Google scholar

 

Original scientific publications in peer-review journals:

 

A95.   Reglero P, E Blanco, A Ortega, Ø Fiksen, F Gándara, M Seoka, FJ Viguri, A Folkvord, 2015. Prey selectivity in piscivorous bluefin tuna larvae reared in the laboratory. Journal of Plankton Research. 37, 2-5.

A94.   Jung K-M, A Folkvord, OS Kjesbu, S Sundby, 2014. Experimental parameterisation of principal physics in buoyancy variations of marine teleost eggs. PlosOne. 9, e104089.

A93.   Frommel AY, RH Maneja, DM Lowe, C Pascoe, AJ Geffen, A Folkvord, U Piatkowski, C Clemmesen, 2014. Organ damage in Atlantic herring larvae as a result of ocean acidification. Ecological Applications. 24, 1131-1143.

A92.   Reglero P, A Ortega, E Blanco, Ø Fiksen, FJ Viguri, F Gándara, M Seoka, A Folkvord, 2014. Size-related differences in growth and survival in piscivorous fish larvae fed different prey types. Aquaculture. 433, 94-101.

A91.   Folkvord A, C Jørgensen, K Korsbrekke, RDM Nash, T Nilsen, JE Skjæraasen, 2014. Trade-offs between growth and reproduction in wild Atlantic cod. Canadian Journal of Fisheries and Aquatic Sciences. 71, 1106-1112.

A90.   Imsland AK, K Ólafsson, S Skírnisdóttir, S Gunnarsson, M Oddgeirsson, S van Damme, SJ Helyar, J Skadal, A Folkvord, 2014. Life history of turbot in Icelandic waters: intra- and inter-population genetic diversity and otolith tracking of environmental temperatures. Fisheries Research. 155, 185-193.

A89.   Vollset KW, IA Catalán, Ø Fiksen, A Folkvord, 2013. Effect of food deprivation on distribution of larval and early juvenile cod in experimental vertical temperature and light gradients. Marine Ecology Progress Series. 475, 191-201.

A88.   Ritchie K, A Folkvord, AK Hufthammer, 2013. Oxygen isotope ratios in cod otoliths used to reveal seasonality of fishing at Late Mesolithic sites in Denmark. Archaeofauna. 22, 95-104.

A87.   Maneja RH, AY Frommel, AJ Geffen, A Folkvord, U Piatkowski, MY Chang, C Clemmesen, 2013. Effects of ocean acidification on the calcification of otoliths of larval Atlantic cod, Gadus morhua L. Marine Ecology Progress Series. 477, 251–258,.

A86.   Hansen TW, A Folkvord, E Grøtan, Ø Sæle, 2013. Genetic ontogeny of pancreatic enzymes in Labrus bergylta larvae and the effect of feed type on enzyme activities and gene expression. Comparative Biochemistry and Physiology Part B: Biochemistry & Molecular Biology. 164, 176-184.

A85.   Karlson S, K Michalsen, A Folkvord, 2013. Age determination of Atlantic halibut (Hippoglossus hippoglossus L.) along the coast of Norway - status, and future potentials. ICES Journal of Marine Science. 70, 50-55.

A84.   Maneja RH, AY Frommel, HI Browman, AB Skiftesvik, C Clemmesen, AJ Geffen, A Folkvord, U Piatkowski, C Durif, R Bjelland, 2013. The swimming kinematics of larval Atlantic cod, Gadus morhua L., are resilient to elevated seawater pCO2. Marine Biology. 160, 1963-1972.

A83.   Jung K-M, A Folkvord, OS Kjesbu, A-L Agnalt, A Thorsen, S Sundby, 2012. Egg buoyancy variability in local populations of Atlantic cod (Gadus morhua). Marine Biology. 159, 1969-1980.

A82.   Ferreira AS, EK Stenevik, KW Vollset, R Korneliussen, A Folkvord, 2012. Vertical migration of Norwegian spring-spawning herring larvae in relation to predator and prey distribution. Marine Biology Research. 8, 605-614.

A81.   Meyer S, EM Caldarone, MA Chicharo, C Clemmesen, AM Faria, C Faulk, A Folkvord, GJ Holt, H Høie, P Kanstinger, A Malzahn, D Moran, C Petereit, JG Støttrup, MA Peck, 2012. On the edge of death: Impacts of temperature, species and body size on RNA-DNA ratios of starving marine fish larvae. Journal of Marine Systems. 93, 11-24.

A80.   Frommel AY, RH Maneja, DM Lowe, A Malzahn, A Geffen, A Folkvord, U Piatkowski, T Reusch, C Clemmesen, 2012. Ocean acidification effects on larvae of a commercially important fish species, Atlantic cod (Gadus morhua). Nature Climate Change. 2, 42-46.

A79.   Koedijk R, AK Imsland, A Folkvord, SO Stefansson, TM Jonassen, A Foss, 2012. Larval rearing environment influences the physiological adaptation in juvenile Atlantic cod, Gadus morhua. Aquaculture International. 20, 467–479.

A78.   Catalán IA, KW Vollset, B Morales-Nin, A Folkvord, 2011. The effect of temperature gradients and stomach fullness on the vertical distribution of larval herring in experimental columns. Journal of Experimental Marine Biology and Ecology. 404, 26-32.

A77.   Geffen AJ, H Høie, A Folkvord, AK Hufthammer, C Andersson, U Ninnemann, RB Pedersen, K Nedreaas, 2011. High-latitude climate variability and its effect on fisheries resources as revealed by fossil cod otoliths. ICES Journal of Marine Science. 68, 1081-1089.

A76.   Vollset KW, A Folkvord, HI Browman, 2011. Foraging behaviour of larval cod (Gadus morhua) at low light intensities. Marine Biology. 158, 1125-1133.

A75.   Imsland AK, R Koedijk, SO Stefansson, A Foss, S Hjorleifsdottir, GO Hreggvidsson, E Otterlei, A Folkvord, 2011. A retrospective approach to fractionize variation in body mass of Atlantic cod Gadus morhua. Journal of Fish Biology. 78, 251-264.

A74.   Meier S, HC Morton, G Nyhammer, BE Grøsvik, V Makhotin, A Geffen, S Boitsov, KA Kvestad, A Bohne-Kjersem, A Goksøyr, A Folkvord, J Klungsøyr, A Svardal, 2010. Development of Atlantic cod (Gadus morhua) exposed to produced water during early life stages. Effects on embryos, larvae, and juvenile fish. Marine Environmental Research. 70, 383-394.

A73.   Folkvord A, R Koedijk, V Lokøy, AK Imsland, 2010. Timing and selectivity of mortality in reared Atlantic cod revealed by otolith analysis. Environmental Biology of Fishes. 89, 513-519.

A72.   Seljeset O, K Vollset, A Folkvord, AJ Geffen, 2010. The role of prey concentration and size range in the growth and survival of larval cod. Marine Biology Research. 6, 251-262.

A71.   Koedijk RM, NR le Francois, PU Blier, A Foss, A Folkvord, D Ditlacadet, SG Lamarre, SO Stefansson, AK Imsland, 2010. Ontogenetic effects of diet during early development on growth performance, myosin mRNA expression and metabolic enzyme activity in Atlantic cod juveniles reared at different salinities. Comparative Biochemistry and Physiology, Part A. 156, 102-109.

A70.   Catalán IA, A Folkvord, I Palomera, G Quilez-Badia, F Kallianoti, A Tselepides, A Kallianotis, 2010. Growth and feeding patterns of European anchovy (Engraulis encrasicolus) early life stages in the Aegean Sea (NE Mediterranean). Estuarine, Coastal and Shelf Science. 86, 299-312.

A69.   Koedijk RM, A Folkvord, A Foss, K Pittman, SO Stefansson, S Handeland, AK Imsland, 2010. The influence of first-feeding diet on the Atlantic cod Gadus morhua phenotype: survival, development and long-term consequences for growth. Journal of Fish Biology. 77, 1-19.

A68.   Hufthammer AK, H Høie, A Folkvord, AJ Geffen, CA Andersson, US Ninnemann, 2010. Seasonality of human site occupation based on stable oxygen isotope ratios of cod otoliths. Journal of Archaeological Science. 37, 78-83.

A67.   Bat NK, CT Vinh, A Folkvord, A Johannessen, K Tsuchiya, S Segawa, 2009. Age and  growth of mitre squid Photololigo chinensis in the Tonkin Gulf of Vietnam based on statolith microstructure. La mer. 47, 57-65.

A66.   Vollset KW, Ø Fiksen, A Folkvord, 2009. Vertical distribution of larval cod (Gadus morhua) in experimental temperature gradients. Journal of Experimental Marine Biology and Ecology. 379, 16-32.

A65.   Folkvord A, Ø Fiksen, H Høie, A Johannessen, E Otterlei, KW Vollset, 2009. What can size distributions within cohorts tell us about ecological processes in fish larvae? Scientia Marina. 74, 119-130.

A64.   Husebø Å, EK Stenevik, A Slotte, P Fossum, A Salthaug, F Vikebø, S Aanes, A Folkvord, 2009. Effects of hatching time on year-class strength in Norwegian spring spawning herring (Clupea harengus). ICES Journal of Marine Science. 66, 1710-1717.

A63.   Folkvord A, H Høie, A Johannessen, T Solbakken, 2009. Effects of prey concentration, light regime and parental origin on growth and survival of herring larvae under controlled experimental conditions. ICES Journal of Marine Science. 66, 1702-1709.

A62.   Vollset KW, O Seljeset, Ø Fiksen, A Folkvord, 2009. A common garden experiment with larval Northeast Arctic and Norwegian coastal cod cohorts in replicated mesocosms. Deep-Sea Research II. 56, 1984-1991.

A61.   Busch KET, A Folkvord, H Otterå, WF Hutchinson, T Svåsand, 2009. Effects of female spawning experience and larval size on feeding and growth of cod larvae Gadus morhua L. reared in mesocosms. Marine Biology Research. 5, 286-296.

A60.   Urtizberea A, Ø Fiksen, A Folkvord, X Irigoien, 2008. Modelling growth of larval anchovies including diel feeding patterns, temperature and body size. Journal of Plankton Research. 30, 1369-1383.

A59.   Li L, H Høie, AJ Geffen, E Heegard, J Skadal, A Folkvord, 2008. Back-calculation of previous fish size using individually tagged and marked Atlantic cod (Gadus morhua). Canadian Journal of Fisheries and Aquatic Sciences. 65, 2496-2508.

A58.   Høie H, A Folkvord, H Mosegaard, L Li, LA Clausen, B Nordberg, AJ Geffen, 2008. Restricted fish feeding reduces cod otolith opacity. Journal of Applied Ichthyology. 24, 138-143.

A57.   Imsland AK, A Foss, R Koedijk, A Folkvord, SO Stefansson, TM Jonassen, 2007. Persistent growth effects of temperature and photoperiod in Atlantic cod Gadus morhua. Journal of Fish Biology. 71, 1371-1382.

A56.   González-Quirós R, I Munuera , A Folkvord, 2007. Cell cycle analysis of brain cells as a growth index in larval cod at different feeding conditions and temperatures. Scientia Marina. 71, 485-497.

A55.   Tian T, Ø Fiksen, A Folkvord, 2007. Estimating larval fish growth under size-dependent mortality: a numerical analysis of bias. Canadian Journal of Fisheries and Aquatic Sciences. 64, 554-562.

A54.   Imsland AK, A Foss, T Alvseike, A Folkvord, SO Stefansson, TM Jonassen, 2007. Interaction between temperature and photoperiod on growth and feeding of Atlantic cod (Gadus morhua): possible secondary effects. Canadian Journal of Fisheries and Aquatic Sciences. 64, 239-248.

A53.   Kristiansen T, Ø Fiksen, A Folkvord, 2007. Modelling feeding, growth, and habitat selection in larval Atlantic cod (Gadus morhua): observations and model predictions in a macrocosm environment. Canadian Journal of Fisheries and Aquatic Sciences. 64, 136-151.

A52.   Bang A, P Grønkjær, A Folkvord, 2007. Possible fitness costs of high and low standard metabolic rates in larval herring Clupea harengus, as determined by otolith microstructure. Marine Ecology Progress Series. 331, 233-242.

A51.   Meisfjord J, F Midtøy, A Folkvord, 2006. Validation of daily increment deposition in otoliths of juvenile Limnothrissa miodon (Clupeidae). Journal of Fish Biology. 69, 1845-1848.

A50.   Imsland AK, A Foss, R Koedijk, A Folkvord, SO Stefansson, TM Jonassen, 2006. Short- and long-term differences in growth, feed conversion efficiency and deformities in juvenile Atlantic cod (Gadus morhua) startfed on rotifers and zooplankton. Aquaculture Research. 37, 1015-1027.

A49.   Caldarone EM, CM Clemmesen, E Berdalet, TJ Miller, A Folkvord, GJ Holt, MP Olivar, IM Suthers, 2006. Intercalibration of four spectrofluorometric protocols for measuring RNA/DNA ratios in larval and juvenile fish. Limnology and Oceanography: Methods. 4, 153-163.

A48.   Simonsen CS, P Munk, A Folkvord, SA Pedersen, 2006. Feeding ecology of Greenland halibut and sandeel larvae off West Greenland. Marine Biology. 149, 937-952.

A47.   Imsland AK, A Foss, A Folkvord, SO Stefansson, TM Jonassen, 2005. The interrelation between temperature regimes and fish size in juvenile Atlantic cod (Gadus morhua): effects on growth and feed conversion efficiency. Fish Physiology and Biochemistry. 31, 347-361.

A46.   Imsland AK, A Foss, A Folkvord, SO Stefansson, TM Jonassen, 2005. Genotypic response to photoperiod treatment in Atlantic cod (Gadus morhua). Aquaculture. 250, 525-532.

A45.   Høie H, A Folkvord, 2006. Estimating the timing of growth rings in Atlantic cod otoliths using stable oxygen isotopes. Journal of Fish Biology. 68, 826-837.

A44.   Imsland AK, A Foss, G Nævdal, T Johansen, A Folkvord, SO Stefansson, TM Jonassen, 2004. Variations in growth in haemoglobin genotypes of Atlantic cod. Fish Physiology and Biochemistry. 30, 47-55.

A43.   Folkvord A, 2005. Comparison of size-at-age of larval Atlantic cod (Gadus morhua) from different populations based on size- and temperature-dependent growth models. Canadian Journal of Fisheries and Aquatic Sciences. 62, 1037-1052.

A42.   Folkvord A, A Johannessen, E Moksness, 2004. Temperature dependent otolith growth in Norwegian spring spawning herring (Clupea harengus) larvae. Sarsia. 89, 297-310.

A41.   Skajaa K, A Fernö, A Folkvord, 2004. Ontogenetic- and condition-related effects of starvation on responsiveness in herring larvae (Clupea harengus L.) during repeated attacks by a model predator. Journal of Experimental Marine Biology and Ecology. 312, 253-269.

A40.   Høie H, CA Dahl, A Folkvord, Ø Karlsen, 2004. Precision and accuracy of stable isotope signals in otoliths of pen reared cod (Gadus morhua L.) when sampled with high resolution micromill. Marine Biology. 144, 1039-1049.

A39.   Høie H, E Otterlei, A Folkvord, 2004. Temperature-dependent fractionation of stable oxygen isotopes in otoliths of juvenile cod (Gadus morhua L). ICES Journal of Marine Science. 61, 243-251.

A38.   Fox CJ, A Folkvord, AJ Geffen, 2003. Otolith micro-increment formation in herring Clupea harengus larvae in relation to growth rate. Marine Ecology Progress Series. 264, 83-94.

A37.   Skajaa K, A Fernö, A Folkvord, 2003. Swimming, feeding and predator avoidance in cod larvae (Gadus morhua L.): trade-offs between hunger and predation risk. in: Browman HI, AB Skiftesvik (Eds.), The Big Fish Bang: Proceedings of the 26th Annual Larval Fish Conference. Institute of Marine Research, Bergen, Norway, pp. 105-121.

A36.   Høie H, A Folkvord, E Otterlei, 2003. Effect of somatic and otolith growth rate on stable isotopic composition of early juvenile cod (Gadus morhua L) otoliths. Journal of Experimental Marine Biology and Ecology. 289, 41-58.

A35.   Tytler P, CJ Fox, A Folkvord, 2002. Glycoconjugates in the otolithic membrane of herring larvae: a possible framework for encoding the life history recorder in fishes. Journal of Fish Biology. 61, 39-49.

A34.   Otterlei E, A Folkvord, G Nyhammer, 2002. Temperature dependent otolith growth of larval and early juvenile Atlantic cod (Gadus morhua). ICES Journal of Marine Science. 59, 851-860.

A33.   Johannessen A, G Blom, A Folkvord, 2000. Differences in growth pattern between spring and autumn spawned herring (Clupea harengus L.) larvae. Sarsia. 85, 461-466.

A32.   Folkvord A, G Blom, A Johannessen, E Moksness, 2000. Growth dependent age estimation in herring (Clupea harengus L.) larvae. Fisheries Research. 46, 91-103.

A31.   Otterlei E, G Nyhammer, A Folkvord, SO Stefansson, 1999. Temperature- and size-dependent growth of larval and early juvenile Atlantic cod (Gadus morhua): a comparative study of Norwegian coastal cod and northeast Arctic cod. Canadian Journal of Fisheries and Aquatic Sciences. 56, 2099-2111.

A30.   Fiksen Ø, A Folkvord, 1999. Modelling growth and ingestion processes in herring Clupea harengus larvae. Marine Ecology Progress Series. 184, 273-289.

A29.   Høie H, A Folkvord, A Johannessen, 1999. The influence of different  parental combinations and incubation temperature on the RNA and DNA content of herring (Clupea harengus L.) larvae at hatching: a pilot study. Journal of Fish Biology. 55, 110-118.

A28.   Høie H, A Folkvord, A Johannessen, 1999. Maternal, paternal and temperature effects on otolith size of young herring (Clupea harengus L.) larvae. Journal of Experimental Marine Biology and Ecology. 234, 167-184.

A27.   Suneetha K-B, A Folkvord, A Johannessen, 1999. Responsiveness of selected condition measures of herring, Clupea harengus, larvae to starvation in relation to ontogeny and temperature. Environmental Biology of Fishes. 54, 191-204.

A26.   Imsland AK, T Nilsen, A Folkvord, 1998. Stochastic simulation of size variation in turbot: possible causes analysed with an individual-based model. Journal of Fish Biology. 53, 237-258.

A25.   Sunde LM, A Imsland, A Folkvord, SO Stefansson, 1998. Effects of size grading on growth and survival of juvenile turbot at two temperatures. Aquaculture International. 6, 19-32.

A24.   Blom G, A Folkvord, A Johannessen, P Fossum, 1997. Interannual variations in condition indices of larval Norwegian spring-spawning herring. Journal of Fish Biology. 51, 370-384.

A23.   Folkvord A, K Rukan, A Johannessen, E Moksness, 1997. Early life history of herring larvae in contrasting feeding environments determined by otolith microstructure analysis. Journal of Fish Biology. 51, 250-263.

A22.   Blom G, A Folkvord, 1997. A snapshot of cannibalism in 0-group Atlantic cod (Gadus morhua) in a marine pond. Journal of Applied Ichthyology-Zeitschrift Fur Angewandte Ichthyologie. 13, 177-181.

A21.   Imsland AK, A Folkvord, ÓDB Jónsdóttir, SO Stefansson, 1997. Effects of exposure to extended photoperiods during the first winter on long-term growth and age at first maturity in turbot (Scophthalmus maximus). Aquaculture. 159, 125-141.

A20.   Bergstad OA, A Folkvord, 1997. Dispersal of tagged juvenile turbot Scophthalmus maximus on the Norwegian Skagerrak coast. Fisheries Research. 29, 211-215.

A19.   Imsland AK, A Folkvord, GL Grung, SO Stefansson, GL Taranger, 1997. Sexual dimorphism in growth and maturation of turbot, Scophthalmus maximus (Rafinesque, 1810). Aquaculture Research. 28, 101-114.

A18.   Folkvord A, L Ystanes, A Johannessen, E Moksness, 1996. RNA:DNA ratios and growth of herring (Clupea harengus L.) larvae reared in mesocosms. Marine Biology. 126, 591-602.

A17.   Imsland AK, LM Sunde, A Folkvord, SO Stefansson, 1996. The interaction of temperature and fish size on growth of juvenile turbot. Journal of Fish Biology. 49, 926-940.

A16.   Folkvord A, IM Haaland, O Grahl-Nielsen, A Johannessen, 1996. Fatty acid composition of herring (Clupea harengus L.) larvae off western Norway. Sarsia. 8, 299-305.

A15.   Stenevik EK, P Fossum, A Johannessen, A Folkvord, 1996. Identification of Norwegian spring spawning herring (Clupea harengus L.) larvae from spawning grounds off western Norway applying otolith microstructure analysis. Sarsia. 80, 285-292.

A14.   Hallaråker H, A Folkvord, SO Stefansson, 1995. Growth of juvenile halibut (Hippoglossus hippoglossus L.) related to temperature, day length and feeding regime. Netherlands Journal of Sea Research. 34, 139-147.

A13.   Imsland AK, A Folkvord, SO Stefansson, 1995. Growth, oxygen consumption and activity of juvenile turbot (Scophthalmus maximus) reared under different temperatures and photo periods. Netherlands Journal of Sea Research. 34, 149-159.

A12.   Folkvord A, E Moksness, 1995. RNA/DNA ratios and growth of herring larvae. Marine Ecology Progress Series. 121, 311-312.

A11.   Folkvord A, V Øiestad, PG Kvenseth, 1994. Growth patterns of three cohorts of Atlantic cod larvae (Gadus morhua L.) studied in a macrocosm. ICES Journal of Marine Science. 51, 325-336.

A10.   Folkvord A, G Blom, O Dragesund, A Johannessen, O Nakken, G Nævdal, 1994. A conceptual framework for enhancing and stabilizing recruitment of marine fish stocks. Aquaculture and Fisheries Management. 25, 245-258.

A09.   Folkvord A, 1993. Prey recognition in stomachs of cannibalistic juvenile cod (Gadus morhua L.). Sarsia. 78, 97-100.

A08.   Otterå H, A Folkvord, 1993. Allometric growth in juvenile cod (Gadus morhua L.) and possible effects on cannibalism. Journal of Fish Biology. 43, 643-645.

A07.   Folkvord A, H Otterå, 1993. Effects of initial size distribution, day length and feeding frequency on growth, survival and cannibalism in juvenile Atlantic cod (Gadus morhua L.). Aquaculture. 114, 243-260.

A06.   Nævdal G, A Folkvord, E Otterlei, S Thorkildsen, 1992. Growth rate related to genotype of 0-group cod at three environmental temperatures. Sarsia. 77, 71-73.

A05.   Booman CI, A Folkvord, JR Hunter, 1991. Responsiveness of starved northern anchovy Engraulis mordax larvae to predatory attacks by adult anchovy. Fishery Bulletin U.S. 89, 707-711.

A04.   Folkvord A, 1991. Growth, survival and cannibalism of cod juveniles (Gadus morhua L.): effects of feed type, starvation and fish size. Aquaculture. 97, 41-59.

A03.   Øiestad V, T Pedersen, A Folkvord, Å Bjordal, PG Kvenseth, 1987. Automatic feeding and harvesting of juvenile Atlantic cod (Gadus morhua L.) in a pond. Modelling Identification and Control. 8, 39-46.

A02.   Folkvord A, JR Hunter, 1986. Size specific vulnerability of northern anchovy (Engraulis mordax) larvae to predation by fishes. Fishery Bulletin U.S. 84, 859-869.

A01.   Øiestad V, PG Kvenseth, A Folkvord, 1985. Mass production of Atlantic cod juveniles (Gadus morhua) in a Norwegian saltwater pond. Transactions of the American Fisheries Society. 114, 590-595.

 

Other scientific publications with original content:

 

B19.   Sundby S, P Fossum, AD Sandvik, FB Vikebø, A Aglen, L Buhl-Mortensen, A Folkvord, K Bakkeplass, P Buhl-Mortensen, M Johannessen, M Jørgensen, T Kristiansen, CS Landa, MS Myksvoll, RDM Nash, 2013. KunnskapsInnhenting Barentshavet – LOfoten – Vesterålen (KILO). Fisken og Havet, 1-188.

B18.   Kristiansen T, Ø Fiksen, A Folkvord, 2005. Larval cod in ponds and models:the role of habitat selection. Globec International Newsletter. 11, 22.

B17.   Belchier M, C Clemmesen, D Cortés, A Doan, A Folkvord, A García, AJ Geffen, H Høie, A Johannessen, E Moksness, Hd Pontual, T Rámirez, D Schnack, B Sveinsbø, 2004. Recruitment studies: Manual of tools for recruitment studies. ICES Techniques in Marine Environmental Science, 1-35.

B16.   Salvanes AGV, A Aksnes, A Folkvord, M Hordnes, H Høie, F Midtøy, J Nejstgaard, E Otterlei, JE Skjæraasen, T Sørnes, AC Utne-Palm, 2003. Environmental influence on fish stocks. IFM Rapport, 1-21.

B15.   Høie H, A Folkvord, A Johannessen, 2000. A multivariate analysis of condition of herring larvae from different environmental conditions, ICES Council Meeting 2000/R:04, pp. 1-10.

B13.   Folkvord A, E Otterlei, A Svardal, 1999. Alkylerte fenolers hormonelle virkning på torsk - generasjonseffekter. Vekst, overlevelse og kjønnsdifferensiering. IFM Rapport, 1-11.

B12.   Fiksen Ø, A Folkvord, 1999. Maternal effects and the benefit of yolk supply in cod larvae in different enfironments - a simulation model, ICES Council Meeting 1999/Y:21, pp. 1-6.

B12.   Fiksen Ø, E Otterlei, A Folkvord, 2000. Experiments and models as reciprocal tools to understand environmental links in recruitment dynamics, ICES Council Meeting 2000/R:02, pp. 1-8.

B11.   Folkvord A, G Blom, JV Helvik, BI Holmqvist, A Johannessen, E Otterlei, SO Stefansson, 1998. Teori- og eksperimentbasert studie av rekruttering hos marin fisk - Sluttrapport fra del I av strategisk program. IFM Rapport, 1-46.

B10.   Folkvord A, A Johannessen, 1998. Otolitt mikrostruktur, presisjon, nøyaktighet og anvendelse. IFM Rapport, 1-19.

B09.   Moksness E, K Rukan, L Ystanes, A Folkvord, A Johannessen, 1995. Comparison of somatic and otolith growth in North Sea herring (Clupea harengus L.) larvae; evaluation of growth dynamics in mesocosms. University of South Carolina Press, Columbia.

B08.   Johannessen A, G Blom, A Folkvord, H Svendsen, 1995. The effect of local wind on the distribution of Norwegian spring spawning herring (Clupea harengus L.) larvae. Elsevier, Amsterdam.

B07.   Folkvord A, 1993. Experimental studies of growth and mortality of larval and juvenile cod (Gadus morhua L.) with special emphasis on cannibalism. University of Bergen, pp. 153.

B06.   Folkvord A, H Otterå, 1991. Effects of size distribution, feeding and light regime on growth and survival of cod juveniles in tanks. European Aquaculture Society, Ghent, Belgium.

B05.   Folkvord A, 1989. Growth and cannibalism of cod fry (Gadus morhua) in intensive systems. European Aquaculture Society, Bredene, Belgium.

B04.   Øiestad V, A Folkvord, PG Kvenseth, 1985. Growth-patterns of Atlantic cod larvae (Gadus morhua L.) from first feeding to metamorphosis studied in a mesocosm, ICES Council Meeting 1985/Mini-Symposium/No. 9.

B03.   Folkvord A, PG Kvenseth, T Pedersen, V Øiestad, 1985. Mass-production of juvenile cod (Gadus morhua L.) in a pond: results and new approaches in 1985, ICES Council Meeting 1985/F:63.

B02.   Folkvord A, 1985. Size specific vulnerability of northern anchovy Engraulis mordax larvae to predation by fishes. University of California, San Diego, pp. 96.

B01.   Øiestad V, PG Kvenseth, A Folkvord, 1983. Mass-production of cod fry in a pond in Western Norway. with additional feeding post-metamorphosis, ICES Council Meeting 1983/F:5, pp. 10.

 

Overview papers, reviews, book chapters, synopsis etc.

 

C06.   Mosegaard H, A Folkvord, PJ Wright, 2002. V. Some uses of individual age data. B. Ecological applications. Ifremer-IRD coedition, Brest, France.

C05.   Folkvord A, H Mosegaard, 2002. V. Some uses of individual age data. A. Age and growth analysis. Ifremer-IRD coedition, Brest, France.

C04.   Wright PJ, J Panfili, A Folkvord, H Mosegaard, J Meunier, 2002. IV. Validation methods. A. Direct validation. Ifremer-IRD coedition, Brest, France.

C03.   Butler JL, A Folkvord, 2000. Section 2: Estimation of fish age and growth. Fisheries Research. 46, 51-52.

C02.   Folkvord A, 1997. Ontogeny of cannibalism in larval and juvenile fish with special emphasis on Atlantic cod, Gadus morhua L. Chapman and Hall Ltd., London.

C01.   Folkvord A, H Botnen, PG Kvenseth, 1991. Erfaringer med oppdrett av torskeyngel i poller i Sør-Norge. Institute of Marine Research, Bergen, Norway.

M.Sc. University of San Diego, USA 1985

Dr. scient. University of Bergen, Norway 1993

Leader of NFR GENSINC project, 2016-2019

Leader of NFR project EMBRC Norway 2016-

Leader of "Herring genetics project" 2013-2015 (ref. L. Andersson, Univ. Uppsala)

Participant in IMR project KILO (Kunnskapsinnhenting li Lofoten og Vesterålen). 2011-2013.

Participant in NFR project “The occurrence of skipped spawning and its importance for population dynamics in Northeast Arctic gadoids”, 2009-2012 (ref. R.D.M. Nash, IMR Bergen)

Participant in UiB-IMR network “Hjort Center for Ecosystem Dynamics“ 2013- (ref. O.S. Kjesbu, IMR Bergen)

Leader of Nordic Marine Academy 2005-2010