Hjemmeside: http://folk.uib.no/nfiaf/
Stilling: professor
Telefon: 55 58 44 56
E-post: Arild.Folkvord@bio.uib.no
Besøksadresse: Thormøhlensgt. 53 A/B
Romnummer: 2G15
The focus of my research is on growth and survival mechanisms in the early life of fishes. The research is experimentally oriented, and covers aspects of marine juvenile production and process oriented recruitment studies.
Size-selective mortality in cultured fish is often due to aggression and cannibalism. A large effort has been made to elucidate factors promoting cannibalism in captive cod (e.g. hunger level, size, size differences etc.). Size- and temperature dependent growth in juvenile flatfish and cod has also been studied with the aim of determining the growth potential under culture conditions. The growth process is very dynamic during the early life stages, and controlled laboratory experiments are carried out to calibrate methods used for growth characterization (e.g. RNA:DNA analysis, otolith microstructure analysis, fatty acid analysis).
The age of field caught fish larvae is often difficult to assess. Fortunately, growth increments are usually formed on a daily basis in the ear stones (otoliths) of fish larvae. It is thus possible to estimate the larval age in days in a similar manner as one determines the age of older fish by counting yearly zones in the otoliths and scales. The growth pattern in the otolith also reflects the previous growth history of the larva, and can be used to determine where, and under what conditions, the larva grew up. The otolith has therefore often been referred to as the "black box" in fish. An important part of the ageing research is related to the accuracy and precision of the age estimation. Knowledge in this field is of great importance in process studies and fisheries management.
fiskelarveøkologi, otolitt analyse, marin yngel produksjon, fiskeri økologi, alder- og vekststudier
Original scientific publications in peer-review journals :
A81. Meyer, S., E. Caldarone, M.A. Chicharo, C. Clemmesen, A. Folkvord, G.J. Holt, H. Høie, A. Malzahn, D. Moran, C. Petereit, J. Støttrup & M.A. Peck. (2012). On the edge of death: Impacts of temperature, species and body size on RNA-DNA ratios of starving marine fish larvae. Journal of Marine Systems. 93: 11-24.
A80. Frommel, A. Y., R.H. Maneja, D.M. Lowe, A. Malzahn, A. Geffen, A. Folkvord, U. Piatkowski, T. Reusch & C. Clemmesen. (2012). Ocean acidification effects on larvae of a commercially important fish species, Atlantic cod (Gadus morhua). Nature Climate Change.2: 42-46.
A79. Koedijk, R., A. Foss, A. Folkvord, S.O. Stefansson, T.M. Jonassen & A.K. Imsland (2012). The influence of first feeding diet on the physiological adaptation of Atlantic cod juveniles (Gadus morhua) to ammonia and oxygen. Aquaculture International. 00: 000-000.
A78. Catalan, I. A., K. W. Vollset, B. Morales-Nin & A. Folkvord. (2011). The effect of temperature gradients and stomach fullness on the vertical distribution of larval herring in experimental columns. Journal of Experimental Marine Biology and Ecology 404: 26-32.
A77. Geffen AJ, Høie H, Folkvord A, Hufthammer AK, Dahl CA, Ninnemann U, Pedersen RB, Nedreaas K (2011) High latitude climate variability and its effect on fishery resources as revealed by fossil otoliths. ICES Journal of Marine Science 68: 1447-1458
A76. Vollset KW, Folkvord A, Browman HI (2011). Foraging behaviour of larval cod (Gadus morhua) at low light intensities Marine Biology 158: 1125-1133
A75. Imsland, A.K., R. Koedijk, A. Foss, S.O. Stefansson, S. Hjörleifsdóttir, G.Ó. Hreggvidsson, E. Otterlei & A. Folkvord (2011). A retrospective approach to fractionize variation in body mass of Atlantic cod Gadus morhua. Journal of Fish Biology 78: 251-264
A74. Meier S, Morton HC, Nyhammer G, Grøsvik BE, Makhotin V, Geffen A, Boitsov S, Kvestad KA, Bohne-Kjersem A, Goksøyr A, Folkvord A, Klungsøyr J, Svardal A (2010) Development of Atlantic cod (Gadus morhua) exposed to produced water during early life stages. Effects on embryos, larvae, and juvenile fish. Marine Environmental Research 70: 383-394
A73. Folkvord A, R. Koedijk, V. Lokøy & A.K. Imsland (2010). Timing and selectivity of mortality in reared Atlantic cod revealed by otolith analysis. Environmental Biology of Fishes, 89: 513-519.
A72. Seljeset, O., K.W. Vollset, A. Folkvord & A.J. Geffen (2010). The role of prey concentration and size range in the growth and survival of larval cod. Marine Biology Research, 6: 251-262.
A71. Koedijk, R., N.R. le Francois, P.U. Blier, A. Foss, A. Folkvord, D. Ditlacadet, S.G. Lamarre, S.O. Stefansson & A.K. Imsland (2010). Ontogenetic effects of diet during early development on growth performance, myosin mRNA expression and metabolic enzyme activity in Atlantic cod juveniles reared at different salinities. Comparative Biochemistry and Physiology Part A, 156: 102-109.
A70. Catalán, I.A., A. Folkvord, I. Palomera, A. Kallianiotis, G. Quílez, A. Tselepides & F. Kallianioti (2010). Inter and intra annual growth and feeding ecology patterns of early life stages of the European anchovy Engraulis encrasicolus in the northeastern Aegean Sea (NE Mediterranean). Estuarine, Coastal and Shelf Science, 86: 299-312.
A69. Koedijk, R., A. Folkvord, A. Foss, K. Pittman, S.O. Stefansson, S. Handeland & A. Imsland (2010). The influence of first feeding diet on the Atlantic cod (Gadus morhua L.) phenotype, with emphasis on growth, survival , RNA:DNA and long term implications. Journal of Fish Biology, 77: 1-19.
A68. Hufthammer, A.K., H. Høie, A. Folkvord, A.J. Geffen, C.A. Andersson, & U.S. Ninnemann (2010). Seasonality of human site occupation based on stable oxygen isotope ratios of cod otoliths. Journal of Archaeological Science, 37: 78-83.
A67. Bat, N.K., C.T. Vinh, A. Folkvord, A. Johannessen, K. Tsuchiya, S. Segawa (2009). Age and growth of mitre squid Photololigo chinensis in the Tonkin Gulf of Vietnam based on statolith microstructure. La mer, 47: 57-65.
A66. Vollset, K.W., Ø. Fiksen & A. Folkvord (2009). Vertical distribution of larval cod (Gadus morhua) in experimental temperature gradients. Journal of Experimental Marine Biology and Ecology, 379: 16-22.
A65. Folkvord, A., Ø. Fiksen, H. Høie, A. Johannessen, E. Otterlei & K.W. Vollset (2009). What can size distributions within cohorts tell us about ecological processes in fish larvae? Scientia Marina, 74 (S1): 119-130.
A64. Husebø, Å., E.K. Stenevik, A. Slotte, P. Fossum, A. Salthaug, F. Vikebø, S. Aanes & A. Folkvord (2009). Effects of hatching time on year-class strength in Norwegian spring spawning herring (Clupea harengus). ICES Journal of Marine Science, 66: 1710-1717.
A63. Folkvord, A., H. Høie, A. Johannessen & T. Solbakken (2009). Effects of prey density, light regime and parental origin on growth and survival of herring larvae under controlled experimental conditions . ICES Journal of Marine Science, 66: 1702-1709.
A62. Vollset, K.W., O. Seljeset, Ø. Fiksen & A. Folkvord (2009). A common garden experiment of larval Northeast Arctic and Norwegian coastal cod cohorts in replicated mesocosms. Deep Sea Research II, 56: 1984-1991.
A61. Busch, K.E.T., A. Folkvord, H. Otterå, W.F. Hutchinson & T. Svåsand (2009). Effects of female spawning experience and larval size on feeding and growth of cod larvae Gadus morhua L. reared in mesocosms. Marine Biology Research, 5: 286-296.
A60. Urtizberea, A., Ø. Fiksen, A. Folkvord & X. Irigoien (2008). Modelling growth of larval anchovies including diel feeding patterns, temperature and body size. Journal of Plankton Research, 30(12): 1369-1383.
A59. Li, L., H. Høie, H., A.J. Geffen, E. Heegaard, J. Skadal & A. Folkvord (2008). Back-calculation of previous fish size based on tagged and marked Atlantic cod (Gadus morhua). Canadian Journal of Fisheries and Aquatic Sciences, 65: 2496-2508.
A58. Høie, H., A. Folkvord, H. Mosegaard, L. Li, L.A. Clausen, B. Nordberg & A.J. Geffen (2008). Reduced feeding influence otolith opacity. Journal of Applied Ichthyology, 24: 138-143.
A57. Imsland, A.K., A. Foss, R. Koedijk, A. Folkvord, S.O. Stefansson & T.M. Jonassen (2007). Persistent growth effects of temperature and photoperiod in Atlantic cod Gadus morhua. Journal of Fish Biology, 71: 1371-1382.
A56. González-Quirós, R., I. Munuera & A. Folkvord (2007). Cell cycle analysis of brain cells as a growth index in larval cod at different feeding conditions and temperatures. Scientia Marina, 71: 485-497.
A55. Tian, T., Ø. Fiksen & A. Folkvord (2007). Estimating larval fish growth under size-dependent mortality: a numerical analysis of bias. Canadian Journal of Fisheries and Aquatic Sciences, 64: 554-562.
A54. Imsland, A.K., A. Foss, T. Alvseike, A. Folkvord, S.O. Stefansson & T.M. Jonassen (2007). Interaction between temperature and photoperiod on growth and feeding of Atlantic cod (Gadus morhua): possible long term effects. Canadian Journal of Aquatic and Fisheries Science, 64: 239-248.
A53. Kristiansen, T., Ø. Fiksen & A. Folkvord (2007). Simulating early life history of pelagic fish larvae: model validation and the role of habitat selection. Canadian Journal of Fisheries and Aquatic Sciences, 64: 136-151.
A52. Bang, A., P. Grønkjær & A. Folkvord (2007). Possible fitness costs of high and low metabolic rates for larval herring Clupea harengus as determined by otolith microstructure. Marine Ecology Progress Series, 331: 233-242.
A51. Meisfjord, J.,F. Midtøy & A. Folkvord (2006). Validation of daily increment deposition rate in otoliths of Limnothrissa miodon (Clupeidae). Journal of Fish Biology, 69: 1845-1848.
A50. Imsland, A. K., A. Foss, R. Koedijk, A. Folkvord, S.O. Stefansson & T.M. Jonassen (2006). Short- and long-term differences in growth, feed conversion efficiency and deformities in juvenile Atlantic cod (Gadus morhua) startfed on rotifers and zooplankton. Aquaculture Research, 37: 1015-1027.
A49. Calderone, E.M., C.M. Clemmesen, E. Berdalet, T.J. Miller, A. Folkvord, G.J. Holt, M.P. Pilar & I.M. Suthers (2006). Intercalibration of four spectrofluorometric methods for measuring RNA-DNA ratios in larval and juvenile fish. Limnology and Oceanography: Methods, 4:153-163.
A48. Simonsen, C.S., P. Munk, A. Folkvord & S.A. Pedersen (2006). Feeding ecology of Greenland halibut and sandeel larvae off West Greenland. Marine Biology, 149: 937-952.
A47. Imsland, A.K., A. Foss, A. Folkvord, S.O. Stefansson, S.O., & T.M. Jonassen (2005). The interrelation between temperature regimes and fish size in juvenile Atlantic cod (Gadus morhua): effects on growth and feed conversion efficiency. Fish Physiology and Biochemistry, 31: 347-361.
A46. Imsland, A.K., A. Foss, A. Folkvord, S.O. Stefansson & T.M. Jonassen (2005). Genotypic response to photoperiod treatment in Atlantic cod (Gadus morhua). Aquaculture, 250: 525-532.
A45. Høie, H. & A. Folkvord (2006). Estimating the timing of growth rings in Atlantic cod otoliths using stable oxygen isotopes. Journal of Fish Biology. 68: 826-837.
A44. Imsland, A.K., A. Foss, G. Nævdal, T. Johansen, A. Folkvord, S.O. Stefansson & T.M. Jonassen (2005). Variations in growth in haemoglobin genotypes of Atlantic cod. Fish Physiology and Biochemistry, 30: 47-55.
A43. Folkvord, A. (2005). Comparison of size-at-age of larval cod (Gadus morhua L.) from different populations based on size- and temperature-dependent models. Canadian Journal of Fisheries and Aquatic Sciences, 62: 1037-1052.
A42. Folkvord, A., A. Johannessen & E. Moksness, E. (2004). Temperature dependent otolith growth in herring (Clupea harengus) larvae. Sarsia, 89: 297-310.
A41. Skajaa, K., A. Fernö & A. Folkvord (2004). Ontogenetic- and condition related effects of starvation on reaction rate in herring larvae (Clupea harengus L.) after repeated attacks with a model predator. Journal of Experimental Marine Biology and Ecology, 312: 253-269.
A40. Høie, H., C.A. Dahl, A. Folkvord & Ø. Karlsen, Ø. (2004). Precision and accuracy of stable isotope signals in otoliths of pen reared cod (Gadus morhua L.) when sampled with high resolution micromill. Marine Biology. 144: 1039-1049.
A39. Høie, H, E. Otterlei & A. Folkvord (2004). Temperature-dependent fractionation of oxygen isotopes in otoliths of juvenile cod (Gadus morhua L). ICES Journal of Marine Science, 61: 243-251.
A38. Fox, C., A. Folkvord, & A.J. Geffen (2003). Otolith micro-increment formation in herring (Clupea harengus L.) larvae growing under high and low growth rates. Marine Ecology-Progress Series, 264: 83-94.
A37. Skajaa K, A. Fernö & A. Folkvord (2003). Swimming, feeding and predator avoidance in cod larvae (Gadus morhua L.): trade-offs between hunger and predation risk. In Browman H, Skiftesvik AB (editors). The Big Fish Bang: Proceedings of the 26th Annual Larval Fish Conference. pp. 105-121.
A36. Høie, H., A. Folkvord & E. Otterlei (2003). Effect of somatic and otolith growth rate on stable isotopic composition of early juvenile cod (Gadus morhua L) otoliths. Journal of Experimental Marine Biology and Ecology, 289: 41-58.
A35. Tytler, P., C. Fox & A. Folkvord (2002). Glycoconjugates in the otolithic membranes of the larvae of herring; a framework for encoding the life history recorder in fish? Journal of Fish Biology, 61: 39-49.
A34. Otterlei, E., A. Folkvord & G. Nyhammer (2002). Temperature dependent otolith growth of larval and early juvenile cod (Gadus morhua). ICES Journal of Marine Science, 59: 401-410.
A33. Johannessen, A., G. Blom, & A. Folkvord (2000). Differences in growth pattern between spring and autumn spawned herring (Clupea harengus L.) larvae. Sarsia, 85: 461-466.
A32. Folkvord, A., G. Blom, A. Johannessen, & E. Moksness (2000). Growth dependent age estimation in herring (Clupea harengus L.) larvae. Fisheries Research, 46: 91-103.
A31. Otterlei, E., G. Nyhammer, A. Folkvord & S.O. Stefansson (1999). Temperature and size dependent growth of larval and juvenile cod (Gadus morhua L.) - a comparative study between Norwegian coastal cod and Northeast Arctic cod. Canadian Journal of Fisheries and Aquatic Sciences, 56: 2099-2111.
A30. Fiksen, Ø. & A. Folkvord (1999). Modelling growth and ingestion processes in herring larvae (Clupea harengus L.). Marine Ecology Progress Series, 184: 273-289.
A29. Høie, H., A. Folkvord & A. Johannessen (1999). The influence of different parental combinations and incubation temperature on the RNA and DNA content of herring (Clupea harengus L.) larvae at hatching: a pilot study. Journal of Fish Biology, 55: 110-118.
A28. Høie, H., A. Folkvord & A. Johannessen, A. (1999). Maternal, paternal and temperature effects on otolith size of young herring (Clupea harengus L.) larvae. Journal of Experimental Marine Biology and Ecology, 234: 167-184.
A27. Suneetha, K.B., A. Folkvord & A. Johannessen (1999). Responsiveness of selected condition measures of herring (Clupea harengus L.) larvae to starvation, in relation to ontogeny and temperature. Environmental Biology of Fishes, 54: 191-204.
A26. Imsland, A.K., T. Nilsen & A. Folkvord (1998). Stochastic simulation of size-variation in turbot: possible causes analysed with an individual-based model. Journal of Fish Biology, 53: 237-258.
A25. Sunde, L.M., A.K. Imsland, A. Folkvord & S.O. Stefansson (1998). Effects of size grading on growth and survival in juvenile turbot (Scophthalmus maximus) reared at two temperatures. Aquaculture International, 6: 19-32.
A24. Blom, G., A. Folkvord, A. Johannessen & P. Fossum (1997). Interannual variations in condition indices of larval Norwegian spring-spawning herring (Clupea harengus). Journal of Fish Biology, 51 (Suppl. A): 370-384.
A23. Folkvord, A., K. Rukan, A. Johannessen & E. Moksness (1997). Early life history of herring (Clupea harengus L.) larvae in contrasting feeding environments determined otolith microstructure analysis. Journal of Fish Biology, 51 (Suppl. A): 250-263.
A22. Blom, G., & A. Folkvord (1997). A snapshot of intracohort cannibalism in juvenile Atlantic cod Gadus morhua in a marine pond. Journal of Applied Ichthyology, 13: 177-182.
A21. Imsland, A.K., A. Folkvord, Jónsdóttir, O.D.B. & S.O. Stefansson (1997). Effect of exposure to different photoperiods during the first winter on long-term growth and age at first maturity in turbot (Scophthalmus maximus Rafinesque). Aquaculture, 159: 125-141.
A20. Bergstad, O.A. & A. Folkvord (1997). Dispersal of tagged juvenile turbot Scophthalmus maximus on the Norwegian Skagerrak coast. Fisheries Research, 29: 211-215.
A19. Imsland, A.K., A. Folkvord, G. Grung, S.O. Stefansson & G.L. Taranger (1997). Sexual dimorphism in growth and maturation of turbot, Scophthalmus maximus Rafinesque. Aquaculture Research, 28: 101-114.
A18. Folkvord, A., L. Ystanes, A. Johannessen & E. Moksness (1996). RNA/DNA ratios and growth of herring (Clupea harengus) larvae reared in mesocosms. Marine Biology, 126: 591-602.
A17. Imsland, A.K., L.M. Sunde, A. Folkvord & S.O. Stefansson (1996). The interaction of temperature and size on growth of juvenile turbot (Scophthalmus maximus Rafinesque) reared under culture conditions. Journal of Fish Biology, 49: 926-940.
A16. Folkvord, A., I.M. Haaland, I.M., O. Grahl-Nielsen & A. Johannessen (1996). Fatty acid composition of herring (Clupea harengus L.) larvae off western Norway. Sarsia, 80: 299-305.
A15. Stenevik, E.K., P. Fossum, A. Johannessen & A. Folkvord (1996). Identification of larvae of Norwegian spring spawning herring from spawning grounds off south-western Norway applying otolith microstructure analysis. Sarsia, 80: 285-292.
A14. Hallaråker, H., A. Folkvord & S.O. Stefansson (1995). Growth of juvenile halibut (Hippoglossus hippoglossus L.) related to temperature, day length and feeding regime. Netherlands Journal of Sea Research, 34: 139-147.
A13. Imsland, A.K., A. Folkvord & S.O. Stefansson (1995). Growth, oxygen consumption and activity of juvenile turbot (Scophthalmus maximus) reared under different temperatures and photoperiods. Netherlands Journal of Sea Research, 34: 149-159.
A12. Folkvord, A. & E. Moksness (1995). RNA/DNA ratios and growth of herring larvae. Marine Ecology Progress Series, 121: 311-312.
A11. Folkvord, A., V. Øiestad & P.G. Kvenseth (1994). Growth-patterns of three cohorts of Atlantic cod larvae (Gadus morhua L.) in a mesocosm. ICES Journal of marine Science, 51: 325-336.
A10. Folkvord, A., G. Blom, O. Dragesund, A. Johannessen, O. Nakken & G. Nævdal (1994). A conceptual framework for enhancing and stabilizing recruitment of marine fish stocks. Aquaculture and Fisheries Management, 25 (suppl. 1): 245-258.
A9. Folkvord, A. (1993). Prey recognition in stomachs of cannibalistic juvenile cod (Gadus morhua L.). Sarsia, 78: 97-100.
A8. Otterå, H. & A. Folkvord (1993). Allometric growth in juvenile cod (Gadus morhua L.) and possible effects on cannibalism. Journal of Fish Biology, 43: 643-645.
A7. Folkvord, A. & H. Otterå (1993). Effects of initial size distribution, day length and feeding frequency on growth, survival and cannibalism in juvenile Atlantic cod (Gadus morhua L.). Aquaculture, 114: 243-260.
A6. Nævdal, G., A. Folkvord, E. Otterlei & S. Torkhildsen (1992). Growth related to genotype of 0-group cod at three environmental temperatures. Sarsia, 77: 71-73.
A5. Booman, C., A. Folkvord & J.R. Hunter (1991). Responsiveness of starved Northern anchovy (Engraulis mordax) larvae to predatory attacks by adult anchovy. Fishery Bulletin U.S., 89: 707-711.
A4. Folkvord, A. (1991). Growth, survival and cannibalism of cod juveniles (Gadus morhua L.): effects of feed type, starvation and fish size. Aquaculture, 97: 41-59.
A3. Øiestad, V., T. Pedersen, A. Folkvord, Å. Bjordal & P.G. Kvenseth (1987). Automatic feeding and harvesting of juvenile Atlantic cod (Gadus morhua L.) in a pond. Modelling Identification and Control, 8(1): 39-46.
A2. Folkvord, A. & J.R. Hunter (1986). Size specific vulnerability of northern anchovy, Engraulis mordax, larvae to predation by fishes. Fishery Bulletin U.S., 84: 859-869.
A1. Øiestad, V., P.G. Kvenseth & A. Folkvord (1985). Mass production of Atlantic cod juveniles Gadus morhua in a Norwegian saltwater pond. Transactions of the American Fisheries Society, 114: 590-595.
Overview papers, reviews, book chapters, synopsis etc.
C6. Mosegaard, H., A. Folkvord & P.J. Wright (2002) Chapter V. Some uses of individual age data. Part B. Ecological applications. In Panfili, J., H. Troadec, H. de Pontual & P.J. Wright (eds). Manual of fish sclerochronology, pp. 167-178. Brest, France. Ifremer-IRD coedition.
C5. Folkvord, A. & H. Mosegaard (2002). Chapter V. Some uses of individual age data. Part A. Age and growth analysis. In Panfili, J., H. Troadec, H. de Pontual & P.J. Wright (eds). Manual of fish sclerochronology, pp. 146-166. Brest, France. Ifremer-IRD coedition.
C4. Wright, P.J., J. Panfili, A. Folkvord, H. Mosegaard & J. Meunier (2002) Chapter IV. Validation methods. Part A. Direct validation. In Panfili, J., H. Troadec, H. de Pontual & P.J. Wright (eds). Manual of fish sclerochronology, pp. 114-127. Brest, France. Ifremer-IRD coedition.
C3. Butler, J.L. & A. Folkvord (2000). Editorial. Section 2: Estimation of fish age and growth. Fisheries Research, 46: 51-52.
C2. Folkvord, A. (1997). Ontogeny of cannibalism in larval and juvenile fish with special emphasis on cod, Gadus morhua L. In: R.C. Chambers and E.D. Trippel (Editors), Early life history and recruitment in fish populations. Chapman and Hall Ltd., London, pp. 251-278.
C1. Folkvord, A., H.B. Botnen & P.G. Kvenseth (1991). Erfaringer med oppdrett av torskeyngel i poller i Sør-Norge. In J.C. Holm and T. Svåsand (eds), Håndbok i torskeoppdrett. pp. 82-95. ISBN 82-7461-025-3. Havforskningsinstituttets Senter for Havbruk.
MAR 338 Fiskelarveøkologi (høst)
BIO 300 Biologisk dataanalyse og forsøksoppsett (høst)
Master program i Fiskeriøkologi og forvaltning (oppgave veiledning)
Master program i Havbruksbiologi (oppgave veiledning)
Master program i Biologi (oppgave veiledning)
Deltaker i HI prosjekt KILO (Kunnskapsinnhenting li Lofoten og Vesterålen). 2011-2013.
Deltaker i NFR prosjekt “The occurrence of skipped spawning and its importance for population dynamics in Northeast Arctic gadoids”, 2009-2012 (ref. R.D.M. Nash, HI Bergen)
Leder for Nordic Marine Academy 2005-2010
Deltaker i UiB-HI nettverk “Johan Hjort Center“ 2008- (ref. S. Sundby, HI Bergen)
Partner in EU network “Understanding the mechanisms of stock recovery” (UNCOVER) 2006-2009 (ref. C. Hammer, Rostock, Germany).
Deltaker i Nordisk Ministerråd Prosjekt, CodLog & MicroCod 2006-2008 (ref. P. Grønkjær, Århus, Danmark)
Deltaker i NFR prosjekt “Progressive, modern production of juvenile Atlantic cod” (PROCOD), 2006-2009 (ref. A.K. Imsland, UiB)