Home
  • E-mailManuel.Malaquias@uib.no
  • Phone+47 55 58 25 82
  • Visitor Address
    Allégaten 41
    Realfagbygget
    5007 Bergen
    Room 
    1C18b
  • Postal Address
    Postboks 7800
    5020 Bergen

How do species originate in the oceans? And what causes their extinction? Why some regions in the oceans are more diverse than others? Are the mechanisms responsible for the patterns of diversity in the deep-sea the same as in shallow ecosystems? Understanding the patterns and causes that drive present diversity in the oceans it’s one of the main goals of my research. As model organisms I use a group of gastropod molluscs – the cephalaspids. The study of the diversity, systematics and evolution of cephalaspids gastropods is consequently an important line of my research activity. This group of marine snails is poorly understood and I use morphological and DNA characters to recognize species and reconstruct hypotheses of evolution  and relationships (phylogenies) at species, genera, and family level.

BIO 232 Systematics and Evolution of Marine Invertebrates (responsible)

Objectives and Content

Obtaining a comprehensive overview of the diversity of invertebrate animals (phyla) and their morphological traits.

Discuss about animal evolution (metazoans) and become familiar with the latest hypotheses of evolutionary relationships based on molecular phylogenetics.

Teaching activities include lectures, seminars and laboratory activities with hands-on anatomical dissections, optical and scanning electron microscopy sessions.

 

 

BIO 299 Research Project in Biology

 

Academic article
  • Show author(s) (2023). Unity in diversity: morphological and genetic variability, integrative systematics, and phylogeography of the widespread nudibranch mollusc Onchidoris muricata. Systematics and Biodiversity.
  • Show author(s) (2023). The genus Pupa Röding, 1798 (Mollusca, Gastropoda, Acteonidae) in New Caledonia with notes on Recent species. Zootaxa. 471-506.
  • Show author(s) (2023). The Pliocene Gastropoda (Mollusca) of Estepona, southern Spain. Part 22: Marine Heterobranchia (excluding Pyramidelloidea). Cainozoic Research. 337-432.
  • Show author(s) (2023). Molecular phylogeny of the marine snail genus Haminoea (Gastropoda, Cephalaspidea): A framework to study marine diversity and speciation. Zoologica Scripta. 52-77.
  • Show author(s) (2023). Integrative taxonomy reveals that not all European reddish runcinids are the same: the case of the Runcina ferruginea Kress, 1977 (Gastropoda, Heterobranchia, Runcinida) species-complex, with the description of a new genus. Invertebrate systematics. 61-77.
  • Show author(s) (2023). FUN Azores: a FUNctional trait database for the meio-, macro-, and megafauna from the Azores Marine Park (Mid-Atlantic Ridge). Frontiers in Ecology and Evolution.
  • Show author(s) (2023). An account on the marine Heterobranchia gastropods collected during the Mission “GAMPA Graciosa 2014” (Archipelago of the Azores, mid-north Atlantic). Arquipélago. Ciências biológicas e marinhas. Life and marine sciences.. 43-49.
  • Show author(s) (2023). A Spanish dancer? No! A troupe of dancers: a review of the family Hexabranchidae Bergh, 1891 (Gastropoda, Heterobranchia, Nudibranchia): A troupe of Spanish dancer. Organisms Diversity & Evolution. 697-742.
  • Show author(s) (2022). Systematic revision of the Indo-West Pacific bubble-snails of the genus Haloa (Pilsbry, 1921) (Cephalaspidea : Haminoeidae). Invertebrate systematics. 436-492.
  • Show author(s) (2022). Polycera norvegica is a valid species, and a plea for good taxonomic practices - A reply to Korshunova et al.2021. Journal of the Marine Biological Association of the United Kingdom.
  • Show author(s) (2022). Neither slugs nor snails: A molecular reappraisal of the gastropod family Velutinidae. Zoological Journal of the Linnean Society. 924-964.
  • Show author(s) (2022). High-level taxonomic splitting in allopatric taxa causes confusion downstream: a revision of the nudibranch family Coryphellidae. Zoological Journal of the Linnean Society. 215-249.
  • Show author(s) (2022). A global phylogeny of the deep-sea gastropod family Scaphandridae (Heterobranchia: Cephalaspidea): Redefinition and generic classification. Molecular Phylogenetics and Evolution.
  • Show author(s) (2021). The taxonomic status of the headshield slug genus Nakamigawaia Kuroda and Habe, 1961 (Gastropoda: Cephalaspidea: Aglajidae), with the description of a new species from the Western Pacific. Journal of Natural History. 2231-2244.
  • Show author(s) (2021). The genus Jorunna (Nudibranchia: Discodorididae) in Europe: A new species and a possible case of incipient speciation. Journal of molluscan studies. 31 pages.
  • Show author(s) (2021). Revisiting the deep-sea Atlantic gastropod species Scaphander gracilis Watson, 1883 (Gastropoda: Cephalaspidea: Scaphandridae): first data on its anatomy, systematics, and ecology. Journal of Natural History. 3053-3066.
  • Show author(s) (2021). Molecular phylogeny of European Runcinida (Gastropoda, Heterobranchia): The discover of an unexpected pool of complex species, with special reference to the case of Runcina coronata. Zoological Journal of the Linnean Society. 761-788.
  • Show author(s) (2021). Molecular and morphological analyses reveal pseudocryptic diversity in Micromelo undatus (Bruguière, 1792) (Gastropoda: Heterobranchia: Aplustridae). Systematics and Biodiversity. 834-858.
  • Show author(s) (2021). Assessment of mitochondrial genomes for heterobranch gastropod phylogenetics. BMC Ecology and Evolution. 1-14.
  • Show author(s) (2021). A molecular phylogeny of Thuridilla Bergh, 1872 sea slugs (Gastropoda, Sacoglossa) reveals a case of flamboyant and cryptic radiation in the marine realm. Cladistics. 647-676.
  • Show author(s) (2020). The temperate Australasian genus Papawera Oskars and Malaquias, 2019 (Gastropoda: Cephalaspidea: Haminoeidae), with a redescription of P. zelandiae and P. maugeansis. Journal of Natural History. 1343-1362.
  • Show author(s) (2020). The copulatory apparatus of the marine gastropod Haminella solitaria (Heterobranchia: Cephalaspidea) and its phylogenetic relevance. Invertebrate biology.. 12 pages.
  • Show author(s) (2020). Systematic revision of the Indo-West Pacific mangrove-associated snails of the genus Bakawan (Cephalaspidea: Haminoeidae). Journal of molluscan studies. 323-341.
  • Show author(s) (2020). Systematic revision of the Indo-West Pacific colourful bubble-snails of the genus Lamprohaminoea Habe, 1952 (Cephalaspidea : Haminoeidae). Invertebrate systematics. 727-756.
  • Show author(s) (2020). Integrative taxonomy reveals a cryptic species of the nudibranch genus Polycera (Polyceridae) in European waters. Journal of the Marine Biological Association of the United Kingdom. 733-752.
  • Show author(s) (2020). A new species of the deep-sea genus Scaphander (Gastropoda, Cephalaspidea) from the Mar del Plata submarine canyon off Argentina. Bulletin of marine science. 111-126.
  • Show author(s) (2020). A massive aggregation of the mangrove snail Bakawan rotundata (A. Adams, 1850) (Cephalaspidea: Haminoeidae) in India. Regional Studies in Marine Science. 1-5.
  • Show author(s) (2019). Unmasking Antarctic mollusc lineages: novel evidence from philinoid snails (Gastropoda: Cephalaspidea). Cladistics. 487-513.
  • Show author(s) (2019). To be or not to be? What molecules say about Runcina brenkoae Thompson, 1980 (Gastropoda: Heterobranchia: Runcinida). Scientia Marina. 223-235.
  • Show author(s) (2019). Beyond shells: first detailed morphological description of the mangrove-associated gastropod Haminoea cf. fusca (A. Adams, 1850) (Cephalaspidea, Haminoeidae), with a COI phylogenetic analysis. Zoosystema. 313-326.
  • Show author(s) (2019). A molecular phylogeny of the gastropod family Haminoeidae sensu lato (Heterobranchia: Cephalaspidea): a generic revision. Invertebrate systematics. 426-472.
  • Show author(s) (2019). A molecular phylogeny of the Indo-West Pacific species of Haloa sensu lato gastropods (Cephalaspidea: Haminoeidae): Tethyan vicariance, generic diversity, and ecological specialization. Molecular Phylogenetics and Evolution. 1-21.
  • Show author(s) (2018). Systematic revision, diversity patterns, and trophic ecology of the tropical Indo-West Pacific sea slug genus Phanerophthalmus A. Adams, 1850 (Cephalaspidea, Haminoeidae). Invertebrate systematics. 1336-1387.
  • Show author(s) (2018). Histological, histochemical, and ultrastructural investigation of the male copulatory apparatus of Haminoea navicula (Gastropoda, Cephalaspidea). Journal of morphology. 554-565.
  • Show author(s) (2018). Diniatys callosa (Preston, 1908), new combination name for Haminoea callosa from the Andaman Islands (India). Journal of Conchology. 213-215.
  • Show author(s) (2018). Description of a new species of Haminoea (Gastropoda: Cephalaspidea) from India, with an account of the diversity of the genus in the Indo-West Pacific. Journal of Natural History. 2437-2456.
  • Show author(s) (2017). Systematic revision of the Antarctic gastropod family Newnesiidae (Heterobranchia: Cephalaspidea) with the description of a new genus and a new abyssal species. Zoological Journal of the Linnean Society. 763-775.
  • Show author(s) (2017). Redescription of the Cephalaspidea gastropod Atys jeffreysi (Weinkauff, 1866) (Haminoeidae), with a discussion on the phylogenetic affinities of the Mediterranean species of the genus. Journal of Natural History. 1593-1608.
  • Show author(s) (2017). Molecular phylogeny of the Aglajidae head-shield sea slugs (Heterobranchia: Cephalaspidea): new evolutionary lineages revealed and proposal of a new classification. Zoological Journal of the Linnean Society. 1-51.
  • Show author(s) (2017). Molecular and morphological systematics of Dolabrifera Gray, 1847 (Mollusca: Gastropoda: Heterobranchia: Aplysiomorpha). Zoological Journal of the Linnean Society. 31-35.
  • Show author(s) (2016). The mediterranean sea as a gateway for invasion of the red sea: The case of the indo-west pacific head-shield slug chelidonura fulvipunctata baba, 1938. Aquatic Invasions. 247-255.
  • Show author(s) (2016). The bubble snails (Gastropoda, Heterobranchia) of Mozambique: an overlooked biodiversity hotspot. Marine Biodiversity. 1-21.
  • Show author(s) (2016). New mediterranean biodiversity records (March 2016). Mediterranean Marine Science. 230-252.
  • Show author(s) (2016). Ecosystem alterations and species range shifts: An Atlantic-Mediterranean cephalaspidean gastropod in an inland Egyptian lake. PLOS ONE.
  • Show author(s) (2016). Diversity and systematics of philinid snails (Gastropoda: Cephalaspidea) in West Africa with remarks on the biogeography of the region. Zoological Journal of the Linnean Society.
  • Show author(s) (2016). Diet preferences of the Aglajidae: a family of cephalaspidean gastropod predators on tropical and temperate shores. Journal of the Marine Biological Association of the United Kingdom. 1101-1112.
  • Show author(s) (2016). A test of color-based taxonomy in nudibranchs: Molecular phylogeny and species delimitation of the Felimida clenchi (Mollusca: Chromodorididae) species complex. Molecular Phylogenetics and Evolution. 215-229.
  • Show author(s) (2015). The opisthobranch gastropods (Mollusca: Heterobranchia) from Venezuela: an annotated and illustrated inventory of species. Zootaxa. 201-256.
  • Show author(s) (2015). Speciation in the dark: Diversification and biogeography of the deep-sea gastropod genus Scaphander in the Atlantic Ocean. Journal of Biogeography. 843-855.
  • Show author(s) (2015). A new phylogeny of the Cephalaspidea (Gastropoda: Heterobranchia) based on expanded taxon sampling and gene markers. Molecular Phylogenetics and Evolution. 130-150.
  • Show author(s) (2014). The family Diaphanidae (Gastropoda: Heterobranchia: Cephalaspidea) in Europe, with a redescription of the enigmatic species Colobocephalus costellatus M. Sars, 1870. Zootaxa. 501-522.
  • Show author(s) (2014). Opisthobranch molluscs of the Azores:results of the IV International Workshop on Malacology and Marine Biology (4–13 July 2011) (Mosteiros, Säo Miguel, Azores). Açoreana : revista de estudos açoreanos. 139-147.
  • Show author(s) (2014). New data on the heterobranch gastropods ('opisthobranchs') for the Bahamas (tropical western Atlantic Ocean). Marine Biodiversity Records. 9 pages.
  • Show author(s) (2014). Diversity and systematics of Haminoeidae gastropods (Heterobranchia: Cephalaspidea) in the tropical West Pacific Ocean: New data on the genera Aliculastrum, Atys, Diniatys and Liloa. Zootaxa. 355-392.
  • Show author(s) (2013). Unique digestive system, trophic specialization, and diversification in the deep-sea gastropod genus Scaphander. Biological Journal of the Linnean Society. 512-525.
  • Show author(s) (2013). Systematic revision of the gastropod family Philinidae (Mollusca: Cephalaspidea) in the northeast Atlantic Ocean with emphasis on the Scandinavian peninsula. Zoological Journal of the Linnean Society. 273-326.
  • Show author(s) (2013). Slipping through the cracks: The taxonomic impediment conceals the origin and dispersal of Haminoea japonica, an invasive species with impacts to human health. PLOS ONE.
  • Show author(s) (2013). New records for the opisthobranch fauna of the Archipelago of the Azores (NE Atlantic Ocean). Marine Biodiversity Records.
  • Show author(s) (2013). New insights into the functional morphology of the male copulatory apparatus of bullid gastropods. Zoomorphology. 145-155.
  • Show author(s) (2013). Epifauna diversity and ecology on intertidal flats in the tropical Niger Delta, with remarks on the gastropod species Haminoea orbignyana. Journal of the Marine Biological Association of the United Kingdom. 249-257.
  • Show author(s) (2012). On the occurrence of the Caribbean sea slug Thuridilla mazda in the eastern Atlantic Ocean. Marine Biodiversity Records.
  • Show author(s) (2011). Marine alien species in Greek Seas: Additions and amendments by 2010. Mediterranean Marine Science. 95-120.
  • Show author(s) (2011). Marine alien species in Greek Seas: Additions and amendments by 2010. Mediterranean Marine Science. 95-120.
  • Show author(s) (2011). Crimora papillata (Nudibranchia: Triophinae), a new record for the shallow marine molluscs of the Azores. Marine Biodiversity Records.
  • Show author(s) (2011). Amphi-Atlantic distributions and cryptic species in sacoglossan sea slugs. Journal of molluscan studies. 401-412.
  • Show author(s) (2010). Systematics, phylogeny, and natural history of Bullacta exarata (Philippi, 1849): an endemic cephalaspidean gastropod from the China Sea. Journal of Natural History. 2015-2019.
  • Show author(s) (2010). Systematics, phylogeny, and natural history of Bullacta exarata (Philippi, 1849): an endemic cephalaspidean gastropod from the China Sea. Journal of Natural History. 2015-2029.
  • Show author(s) (2009). Reassessment of the trophic position of Bullidae (Gastropoda: Cephalaspidea) and the importance of diet in the evolution of cephalaspidean gastropods. Journal of Zoology. 88-97.
  • Show author(s) (2009). Molluscan diversity in the North Atlantic Ocean: New records of opisthobranchs gastropods from the Archipelago of the Azores. Marine Biodiversity Records. e38.
  • Show author(s) (2009). Aplysiopsenes: an additional example of marine polyketides with a mixed acetate/propionate pathway. Tetrahedron Letters. 527-529.
  • Show author(s) (2009). Aplysiopsenes: an additional example of marine polyketides with a mixed acetate/propionate pathway. Tetrahedron Letters. 527-529.
  • Show author(s) (2009). A molecular phylogeny of the Cephalaspidea sensu lato (Gastropoda: Euthyneura): Architectibranchia redefined and Runcinacea reinstated. Zoologica Scripta. 23-41.
  • Show author(s) (2008). Systematic revision of the Recent species of Bullidae (Mollusca: Gastropoda: Cephalaspidea), with a molecular phylogenetic analysis. Zoological Journal of the Linnean Society. 453-543.
  • Show author(s) (2008). Molluscan diversity in the North Atlantic Ocean: New records of opisthobranchs gastropods from the Archipelago of the Azores. Journal of the Marine Biological Association of the United Kingdom. 1-10.
  • Show author(s) (2008). Functional morphology of the gonoduct of the gastropod Bulla striata Bruguière, 1792 (Opisthobranchia: Cephalaspidea): evidence for a monaulic system. Acta Zoologica. 205-210.
Lecture
  • Show author(s) (2016). On the origin and biogeography of the sap-sucking sea slugs genus Thuridilla.
  • Show author(s) (2016). Diversity, systematics, and trophic ecology of the tropical Indo-West Pacific genus Phanerophthalmus (Cephalaspidea, Haminoeidae).
  • Show author(s) (2015). New challenges in Cephalaspidea systematic.
  • Show author(s) (2015). Broad sampling and molecular phylogenetics reveals rampant paraphyly in Haminoeidae cephalaspids: the epilogue.
Academic lecture
  • Show author(s) (2016). Fine scale geographic diversification in the tropical western Atlantic: the phylogeography of Bulla occidentalis.
  • Show author(s) (2015). Bichos estranhos: sistemática e biologia de alguns filos menos conhecidos do Reino Animal.
Editorial
  • Show author(s) (2011). Studies on opisthobranch molluscs: Proceedings of a symposium, World Congress of Malacology, Phuket, Thailand, July 2010. Introduction. Journal of molluscan studies. 321-321.
Popular scientific article
  • Show author(s) (2018). Innsamling av marine virvelløse dyr. Årbok for Universitetsmuseet i Bergen. 101-111.
  • Show author(s) (2009). Darwin: polvos, vinagreiras e fósseis na terra da morabeza [Darwin: octopuses, sea hares, and fossils in the land of “morabeza”]. Portugala. 3-4.
  • Show author(s) (2009). Bermudas: no triângulo à procura de opistobrânquios [Bermuda: seeking opisthobranchs in the triangle]. Portugala. 47-49.
Letter to the editor
  • Show author(s) (2017). The Suez Canal as a revolving door for marine species: A reply to Galil et al. (2016). Aquatic Invasions. 1-4.
Doctoral dissertation
  • Show author(s) (2019). The systematics and phylogeny of Haminoeidae (Mollusca, Cephalaspidea), with special focus on the taxonomy and phylogeny of the genus Haloa sensu lato.
  • Show author(s) (2018). Evolutionary history, connectivity and habitat-use of annelids from deep-sea chemosynthesis-based ecosystems, with an emphasis on the Arctic mid-Ocean Ridge and the Nordic Seas.
  • Show author(s) (2017). The sea slugs of the family Aglajidae (Gastropoda: Cephalaspidea). Dietary specialization and molecular phylogeny of the family, with species delimitation analysis and biogeography of two genera from the clade Chelidonura sensu lato.
Academic chapter/article/Conference paper
  • Show author(s) (2011). Family Haminoeidae Pilsbry, 1895 – Review and commented. 1 pages.
  • Show author(s) (2011). Family Cylichnidae H. Adams & A. Adams, 1854 – Review and commented. 1 pages.
  • Show author(s) (2011). Family Bullinidae J. E. Gray, 1850 – Review and commented. 1 pages.
  • Show author(s) (2011). Family Bullidae d’Orbigny, 1827 – Review and commented. 1 pages.
  • Show author(s) (2011). Family Acteonidae d’Orbigny, 1842 – Review and commented. 1 pages.
  • Show author(s) (2011). Familia Haminoeidae. 5 pages.
  • Show author(s) (2011). Familia Bullidae. 2 pages.
Encyclopedia article
  • Show author(s) (2023). Euopisthobranchs: Skeletal/Support System. 1-7. In:
    • Show author(s) (2023). Microscopic Anatomy of Animals.
  • Show author(s) (2023). Euopisthobranchs: Respiratory System. 1-8. In:
    • Show author(s) (2023). Microscopic Anatomy of Animals.
  • Show author(s) (2023). Euopisthobranchs: Introduction. 1-9. In:
    • Show author(s) (2023). Microscopic Anatomy of Animals.
  • Show author(s) (2023). Euopisthobranchs: Intestinal System. 1-24. In:
    • Show author(s) (2023). Microscopic Anatomy of Animals.
  • Show author(s) (2023). Euopisthobranchs: Excretory System. 1-11. In:
    • Show author(s) (2023). Microscopic Anatomy of Animals.
  • Show author(s) (2023). Euopisthobranchs: Circulatory System. 1-9. In:
    • Show author(s) (2023). Microscopic Anatomy of Animals.
  • Show author(s) (2023). Euopisthobranchs: Body Cavities. 1-10. In:
    • Show author(s) (2023). Microscopic Anatomy of Animals.
Museum exhibition
  • Show author(s) (2019). The whale hall - The great swarm.
Abstract
  • Show author(s) (2017). Investigating the marine invertebrate fauna of the West African continental shelf with DNA barcodes . Genome. 1010-1011.
  • Show author(s) (2010). Complex evolution in the Neotropics: Understanding Caribbean marine biogeography using Bulla occidentalis as a model species. Tropical Natural History. 267.
Poster
  • Show author(s) (2020). Two centuries of marine biodiversity collections and research.
  • Show author(s) (2017). Investigating the marine invertebrate fauna of the West African continental shelf with DNA barcodes.
  • Show author(s) (2016). Marin biodiversitetsforskning ved Universitetsmuseet i Bergen.
  • Show author(s) (2016). Diversity and systematics of the Indo-West Pacific genus Haloa (Euopisthobranchia, Cephalaspidea).
Academic literature review
  • Show author(s) (2017). Haminoea orteai Talavera, Murillo & Templado, 1987 (Mollusca: Gastropoda: Heterobranchia: Cephalaspidea), a widespread species in the Mediterranean and northeastern Atlantic. CBM - Cahiers de Biologie Marine. 107-113.
  • Show author(s) (2013). Systematic revision of the genus Scaphander (Gastropoda, Cephalaspidea) in the Atlantic Ocean, with a molecular phylogenetic hypothesis. Zoological Journal of the Linnean Society. 389-429.
  • Show author(s) (2013). Systematic revision of the gastropod family Philinidae (Mollusca: Cephalaspidea) in the north-east Atlantic Ocean with emphasis on the Scandinavian Peninsula. Zoological Journal of the Linnean Society. 273-326.
  • Show author(s) (2009). Tethyan vicariance, reliction and speciation: evidence from a global molecular phylogeny of the opisthobranch genus Bulla. Journal of Biogeography. 1760-1777.

More information in national current research information system (CRIStin)

122. Lobo-da-Cunha, A., Bartolomaeus, T. and Malaquias, M. A. E. 2023. Euopisthobranchs. Microscopic Anatomy of Animals, in press.

121. Turani, M., Valdés, Á., and Malaquias, M. A. E. 2023. Molecular phylogeny of the marine snail genus Haminoea (Gastropoda, Cephalaspidea): a framework to study marine diversity and speciation. Zoologica Scripta, https://doi.org/10.1111/zsc.12627.

120. Ekimova, I. A., Nikitenko, E., Stanovova, M. V., Schepetov, D. M., Antokhina, T. I., Malaquias, M. A. E., and Valdés, Á. 2023. Unity in diversity: morphological and genetic variability, integrative systematics, and phylogeography of the widespread nudibranch mollusc Onchidoris muricataSystematics and Biodiversity, 21:1, https://DOI: 10.1080/14772000.2023.2246472

119. Tibiriçá, Y., Pola, M., Pittman, C., Gosliner, T. M., Malaquias, M. A., and Cervera, J. L. 2023. A Spanish dancer? No! A troupe of dancers: a review of the family Hexabranchidae Bergh, 1891 (Gastropoda, Heterobranchia, Nudibranchia). Organisms Diversity & Evolutionhttps://doi.org/10.1007/s13127-023-00611-0

118. Malaquias, M. A. E. and Afonso, P. 2023. An account on the marine Heterobranchia gastropods collected during the Mission “GAMPA Graciosa 2014” (Archipelago of the Azores, mid-north Atlantic). Arquipelago. Life and Marine Sciences, 38: 43–49. https://doi.org/10.25752/arq.31347 

117. Giacomello, E., Catarino, D., Gooday, A. J., Berning, B., Figuerola, B., Malaquias, M. A. E., Moura, C. J., Xavier, J. R., Sutton, T. T., Fauconnet, L., Ramalho, S. P., Neves, B. d. M., Machado, G. M., Horton, T., Gebruk, A.V., Minin, K., Bried, J., Molodtsova, T., Silva, M. A., Dilman, A., Kremenetskaia, A., Costa, E. F. S., Clarke, J., Martins, H. R., Pham, C. K., Carreiro-Silva, M., and Colaço, A. 2023. FUN Azores: a FUNctional trait database for the meio-, macro-, and megafauna from the Azores Marine Park (Mid-Atlantic Ridge). Frontiers in Ecology and Evolution, 11: 1050268. doi: 10.3389/fevo.2023.1050268 

116. Valdés, Á., Feliciano, K., and Malaquias, M. A. 2023. The genus Pupa Röding, 1798 (Mollusca, Gastropoda, Acteonidae) in New Caledonia with notes on Recent species. Zootaxa, 5270(3), 471–506. 10.11646/ZOOTAXA.5270.3.4

115. Araújo, A. K., Pola, M., Malaquias, M. A. E., Vitale, F., and Cervera, J. L. 2023. Integrative taxonomy reveals that not all European reddish runcinids are the same: the case of the Runcina ferruginea Kress, 1977 (Gastropoda, Heterobranchia, Runcinida) species-complex, with the description of a new genus. Invertebrate Systematics, 37(1), 61–77. https://doi.org/10.1071/IS22014

114. Fassio, G., Stefani, M., Russini, V., Buge, B., Bouchet, P., Treneman, N., Malaquias, M. A. E., Schiaparelli, S., Modica, M. V., Oliverio, M. 2023. Neither slugs nor snails: a molecular reappraisal of the gastropod family Velutinidae, Zoological Journal of the Linnean Society, 197(4), 924–964. https://doi.org/10.1093/zoolinnean/zlac091

113. Oskars, T. R., and Malaquias, M. A. E. 2022. Systematic revision of the Indo-West Pacific bubble-snails of the genus Haloa (Pilsbry, 1921) (Cephalaspidea: Haminoeidae). Invertebrate Systematics, 36(5): 436–492. https://doi.org/10.1071/IS21011

112. Ekimova, I., Valdés, Á., Malaquias, M. A. E., Rauch, C., Chichvarkhin, A., Mikhlina, A., Antokhina, T., Chichvarkhina, O., and Schepetov, D. 2022. High-level taxonomic splitting in allopatric taxa causes confusion downstream: a revision of the nudibranch family Сoryphellidae. Zoological Journal of the Linnean Society, 196, 215–249. https://doi.org/10.1093/zoolinnean/zlab109

111. Siegwald, J., Kano, Y., and Malaquias, M. A. E. 2022. A global phylogeny of the deep-sea gastropod family Scaphandridae (Heterobranchia: Cepahalaspidea): redefinition and generic classification. Molecular Phylogenetics and Evolution, 169: 107415. https://doi.org/10.1016/j.ympev.2022.107415

110. Hellem, E., and Malaquias, M. A. E. 2021. The taxonomic status of the headshield slug genus Nakamigawaia Kuroda and Habe, 1961 (Gastropoda: Cephalaspidea: Aglajidae), with the description of a new species from the Western Pacific. Journal of Natural History, 55(35–36): 2231–2244. https://doi.org/10.1080/00222933.2021.1986165

109. Siegwald, J. and Malaquias, M. A. E. 2021. Revisiting the deep-sea Atlantic gastropod species Scaphander gracilisWatson, 1883: First data on its morphology, systematics, and ecology. Journal of Natural History, 55(47–48): 3053–3066. https://doi.org/10.1080/00222933.2022.2036378

108. Neuhaus, J., Rauch, C., Bakken, T., Picton, B., Pola, M. and Malaquias, M. A. E. 2021. The genus Jorunna (Nudibranchia: Discodorididae) in Europe: a new species and a possible case of incipient speciation. Journal of Molluscan Studies, 87(4), eyab028. DOI: 10.1093/mollus/eyab028

107. Feliciano, K., Malaquias, M. A. E., Stout, C., Brenzinger, B., Gosliner, T. M., and Valdés, Á. 2021. Molecular and morphological analyses reveal pseudocryptic diversity in Micromelo undatus  (Bruguière, 1792) (Gastropoda: Heterobranchia: Aplustridae). Systematics and Biodiversity, 19(7): 834–858. https://dx.doi.org/10.1080/14772000.2021.1939458

106. Araújo, A. K., Pola, M., Malaquias, M. A. E., Ballesteros, M., and Cervera, J. L. 2021. Molecular phylogeny of European Runcinida (Gastropoda, Heterobranchia): the discovery of an unexpected pool of species complex, with special reference to the case of Runcina coronata (Quatrefages, 1844). Zoological Journal of the Linnean Society, 194: 761–788. https://doi.org/10.1093/zoolinnean/zlab041

105. Martín-Hervás, M. R., Carmona, L., Malaquias, M. A. E., Krug, P. J., Gosliner, T. M. and Cervera, J. L. 2021. A molecular phylogeny of ThuridillaBergh, 1872 sea slugs (Gastropoda, Sacoglossa) reveals a case of flamboyant and cryptic radiation in the marine realm. Cladistics, 37: 647–676. 

104. Varney, R. M., Brenzinger, B., Malaquias, M. A. E., Meyer, C. P., Schrödl, M. and Kocot, K. 2021.  Assessment of mitochondrial genomes for heterobranch gastropod phylogenetics. BMC Ecology and Evolution, 21(1): 1-14. https://doi.org/10.1186/s12862-020-01728-y

103. Bharate, M., Apte, D., and Malaquias, M. A. E. 2020. A massive aggregation of the mangrove snail Bakawan rotundata (A. Adams, 1850) (Cephalaspidea: Haminoeidae) in India. Regional Studies in Marine Science, 39: 101406. https://doi.org/10.1016/j.rsma.2020.101406

102. Sørensen, C. G., Rauch, C., Pola, M. and Malaquias, M. A. E. 2020. Integrative taxonomy reveals a cryptic species of the nudibranch genus Polycera (Polyceridae) in European waters. Journal of the Marine Biological Association of the United Kingdom, 100(5): 733-752. https://doi.org/10.1017/S0025315420000612.

101. Oskars, T. R., and Malaquias, M. A. E. 2020. The temperate Australasian genus Papawera Oskars and Malaquias, 2019 (Gastropoda: Cephalaspidea: Haminoeidae), with a redescription of P. zelandiae and P. maugeansisJournal of Natural History, 54(21-22): 1343-1362.  https://doi.org/10.1080/00222933.2020.1791996.

100. Oskars, T. R., and Malaquias, M. A. E. 2020. Systematic revision of the Indo-West Pacific colourful bubble-snails of the genus Lamprohaminoea Habe, 1952 (Cephalaspidea : Haminoeidae). Invertebrate Systematics, 34: 727–756. https://doi.org/10.1071/IS20026.

99. Oskars, T. R., and Malaquias, M. A. E. 2020. Systematic revision of the Indo-West Pacific mangrove-associated snails of the genus Bakawan (Cephalaspidea: Haminoeidae). Journal of Molluscan Studies, 86: 323–341. doi:10.1093/mollus/eyaa012

98. Lobo-da-Cunha, A., Alves, Â., Oliveira, E., Wranik, W., and Malaquias, M. A. E. 2020. The copulatory apparatus of the marine gastropod Haminella solitaria (Say, 1822) (Heterobranchia: Cephalaspidea) and its phylogenetic relevance. Invertebrate Biology, 139(1): e12281. https://doi.org/10.1111/ivb.12281

97. Siegwald, J., Pastorino, G., Oskars, T., and Malaquias, M. A. E. 2020. A new species of the deep-sea genus Scaphander (Gastropoda, Cephalaspidea) from the Mar del Plata submarine canyon off Argentina. Bulletin of Marine Sciences, 96(1):111–126. https://doi.org/10.5343/bms.2019.0069

96. Oskars, T. R. and Malaquias, M. A. E. 2019. A molecular phylogeny of the Indo-West Pacific species of Haloa sensu lato gastropods (Cephalaspidea: Haminoeidae): Tethyan vicariance, generic diversity, and ecological specialization. Molecular Phylogenetics and Evolution, 139:106557. https://doi.org/10.1016/j.ympev.2019.106557

95. Araujo, A. K., Pola, M., Malaquias, M. A. E., Cervera, J. L. 2019. To be or not to be? What molecules say about Runcina brenkoae Thompson, 1980 (Gastropoda: Heterobranchia: Runcinida). Sciencia Marina, 83(3): 223-235. https://doi.org/10.3989/scimar.04907.07A

94. Aslam, S., Oskars, T. R., Siddiqui, G., Malaquias, M. A. E. 2019. Beyond shells: First detailed morphological description of the mangrove-associated gastropod Haminoea cf. fusca (A. Adams, 1850) (Cephalaspidea: Haminoeidae), with a COI phylogenetic analysis. Zoosystema, 41 (16): 313-326. https://doi.org/10.5252/zoosystema2019v41a16

93. Moles, J., Avila, C., Malaquias, M. A. E. 2019. Unmasking Antarctic mollusc lineages: novel evidence from philinoid snails (Gastropoda: Cephalaspidea). Cladistics, 35: 487–513. https://doi.org/10.1111/cla.12364

92. Oskars, T. R., Too, C. C., Rees, D., Mikkelsen, P., Willassen, E., Malaquias, M. A. E. 2019. A molecular phylogeny of the gastropod family Haminoeidae sensu lato (Heterobranchia, Cephalaspidea), - a generic revision. Invertebrate Systematics, 33(2): 426-472.

91. Bharate, M., Oskars, T. R., Narayana, S., Ravinesh, R., Kumar, A. B., Malaquias, M. A. E. 2018. Description of a new species of Haminoea (Gastropoda: Cephalaspidea) from India, with an account of the diversity of the genus in the Indo-West Pacific. Journal of Natural History, 52 (37–38): 2437–2456. https://doi.org/10.1080/00222933.2018.1533598

90. Austin, J., Gosliner, T., Malaquias, M. A. E. 2018. Systematic revision, diversity patterns, and trophic ecology of the tropical Indo-West Pacific sea slug genus Phanerophthalmus A. Adams, 1850 (Cephalaspidea, Haminoeidae). Invertebrate Systematics, 32: 1336–1387. https://doi.org/10.1071/IS17086

89. Valdés, Á., Breslau, E., Padula, V., Schrödl, M., Camacho, Y., Malaquias, M. A. E., Alexander, J., Bottomley, M., Vital, X., and Gosliner, T. M. 2018. Molecular and morphological systematics of Dolabrifera Gray, 1847 (Mollusca, Gastropoda, Heterobranchia, Aplysiomorpha). Zoological Journal of the Linnean Society, 184: 31–65. https://doi.org/10.1093/zoolinnean/zlx099

88. Moles, J., Ávila, C. and Malaquias, M. A. E. 2018. Systematic revision of the Antarctic gastropod family Newnesiidae (Heterobranchia: Cephalaspidea) with the description of a new genus and a new abyssal species. Zoological Journal of the Linnean Society, 183: 763–775. https://doi.org/10.1093/zoolinnean/zlx089

87. Bharate, M., Oskars, T, Malaquias, M. A. E. 2018. Diniatys callosa (Preston, 1908), new combination name for Haminoea callosa from the Andaman Islands (India). Journal of Conchology, 43(1): 1–3.

86. Lobo-da-Cunha, A., Alves, Â., Oliveira, E. and Malaquias, M. 2018. Histological, histochemical and ultrastructural investigation of the male copulatory apparatus of Haminoea navicula (Gastropoda, Cephalaspidea). Journal of Morphology, 279(4): 554–565. https://doi.org/10.1002/jmor.20790 

85. Zamora-Silva, A. & Malaquias, M. A. E. 2018. Molecular phylogeny of the Aglajidae head-shield sea slugs (Heterobranchia: Cephalaspidea): new evolutionary lineages revealed and proposal of a new classification. Zoological Journal of the Linnean Society, 183(1): 1–51. https://doi.org/10.1093/zoolinnean/zlx064

84. Oskars, T. R., Mifsud, C., & Malaquias, M. A. E. 2017. Redescription of the Cephalaspidea gastropod Atys jeffreysi (Weinkauff, 1866) (Haminoeidae), with a discussion on the phylogenetic affinities of the Mediterranean species of the genus. Journal of Natural History, 51: 1593–1608. https://doi.org/10.1080/00222933.2017.1340526

83. Tibiriça, Y. & Malaquias, M. A. E. 2017. The bubble snails (Gastropoda, Heterobranchia) of Mozambique: an overlooked biodiversity hotspot. Marine Biodiversity, 47(3), 791-811. https://doi.org/10.1007/s1252

82. Malaquias, M. A. E., Zamora-Silva, A., Vitale, D., Spinelli. A., De Matteo, S., Giacobbe, S., Ortigosa, D. & Cervera J. L. 2017. The Suez Canal as a revolving door for marine species: a reply to Galil et al. (2016). Aquatic Invasions, 12(1): 1–4. DOI: https://doi.org/10.3391/ai.2017.12.1.01

81. Garabedian, K., Malaquias, M.A.E., Crocetta, F., Zenetos, A., Kavadas, S. & Valdés, Á. 2017. Haminoea orteai Talavera, Murillo and Templado, 1987 (Mollusca: Gastropoda: Heterobranchia: Cephalaspidea), a widespread species in the Mediterranean and northeastern Atlantic. Cahiers de Biologie Marine, 58(1): 107-113. DOI: 10.21411/CBM.A.19015AD4

80. Malaquias, M. A. E., Ohnheiser, L. T., Oskars, T. R. & Willassen, E. 2017. Diversity and systematics of philinid snails (Gastropoda: Cephalaspidea) in West Africa with remarks on the biogeography of the region. Zoological Journal of the Linnean Society, 180(1): 1-35. https://doi.org/10.1111/zoj.12478

79. Padula, V., Bahia, J., Stöger, I., Camacho-García, Y., Malaquias, M. A. E., Cervera, J. L., & Schrödl, M. 2016. A test of color-based taxonomy in nudibranchs: molecular phylogeny and species delimitation of the Felimida clenchi (Mollusca: Chromodorididae) species complex. Molecular Phylogenetics and Evolution, 103: 215–229. doi:10.1016/j.ympev.2016.07.019

78. Cruz-Rivera, E. & Malaquias, M. A. E. 2016. Ecosystem alterations and species range shifts: An Atlantic-Mediterranean cephalaspidean gastropod in an inland Egyptian lake. PLoS ONE, 11(6): e0156760. doi:10.1371/journal.pone.0156760.

77. Malaquias, M. A. E., Zamora-Silva, A., Vitale, D., Spinelli. A., De Matteo, S., Giacobbe, S., Ortigosa, D. & Cervera J. L. 2016. The Mediterranean Sea as a gateway for invasion of the Red Sea: The case of the Indo-West Pacific head-shield slug Chelidonura fulvipunctata Baba, 1938. Aquatic Invasions, 11(3): 247–255. 

76. Malaquias, M. A. E, Madrenas, E, & Ballesteros, M. 2016. First occurrence of the tropical Indo-West Pacific head-shield sea slug Chelidonura fulvipunctata in the Balearic Islands confirms its range extension into the western Mediterranean Sea. Mediterranean Marine Science 17 (New Mediterranean Biodiversity Records, March 2016): 232.

75. Zamora, B. A. & Malaquias, M. A. E. 2016. Diet preferences of the Aglajidae: a family of cephalaspidean gastropod predators on tropical and temperate shores. Journal of the Marine Biological Association of the United Kingdom, 96(5): 1101–1112. http://doi:10.1017/S0025315415000739

74. Caballer, M., Ortea J., Rivero, N., Carias, G., Malaquias, M. A. E. & Narciso, S. 2015. The opisthobranch gastropods (Mollusca: Heterobranchia) from Venezuela: an annotated and illustrated inventory of species. Zootaxa, 4034: 201–256. 

73. Oskars, T. R., Bouchet, P., & Malaquias, M. A. E. 2015. A new phylogeny of the Cephalaspidea (Gastropoda: Heterobranchia) based on expanded taxon sampling and gene markers. Molecular Phylogenetics and Evolution, 89: 130–150. 

72. Eilertsen, M. E. & Malaquias, M. A. E. 2015. Speciation in the dark: diversification and biogeography of the deep-sea gastropod genus Scaphander in the Atlantic Ocean. Journal of Biogeography, 42: 843–855.

71. Malaquias, M. A. E., Calado, G., Cruz, J. F., and Jensen, K. 2014. Opisthobranch molluscs of the Azores:results of the IV International Workshop on Malacology and Marine Biology (4–13 July 2011) (Mosteiros, Säo Miguel, Azores). Açoreana, Suplemento 10: 139-147.

70. Too, C. C, Carlson, C., Hoff, P. J. and Malaquias, M. A. E. 2014. Diversity and systematics of Haminoeidae gastropods (Heterobranchia: Cephalaspidea) in the tropical West Pacific: new data on the genera AliculastrumAtysDiniatys and LiloaZootaxa, 3794(3): 355-392.

69. Malaquias, M.A.E. 2014. New data on the heterobranch gastropods (“opisthobranchs”) for the Bahamas (tropical western Atlantic ocean). Marine Biodiversity Records, 7, e27.

68. Ohnheiser, L. T. & Malaquias, M. A. E. 2014. The family Diaphanidae (Gastropoda: Heterobranchia: Cephalaspidea) in Europe, with a redescription of the enigmatic species Colobocephalus costellatus M. Sars, 1870. Zootaxa, 3774(6): 501-522.

67. Hanson, D., Cooke, S., Hirano, Y., Malaquias, M. A. E., Crocetta, F., & Valdés, Á. 2013. Slipping through the cracks: The taxonomic impediment conceals the origin and dispersal of Haminoea japonica, an invasive species with impacts to human health. PLoS ONE, 8(10): e77457.

66. Eilertsen, M. E. & Malaquias, M. A. E. 2013. Unique digestive system, trophic specialization, and diversification in the deep-sea gastropod genus ScaphanderBiological Journal of the Linnean Society, 109(3): 512-525..

65. Amorim, A., Oliveira, E., Malaquias, M. A. E. & Lobo-da-Cunha, A. 2013. New insights into the functional morphology of the male copulatory apparatus of bullid gastropods. Zoomorphology, 132: 145-155.

64. Eilertsen, M. E. & Malaquias, M. A. E. 2013. Systematic revision of the genusScaphander (Gastropoda, Cephalaspidea) in the Atlantic Ocean with a molecular phylogenetic hypothesis. Zoological Journal of the Linnean Society, 167(3): 389-429.

63. Cordeiro, R., Malaquias, M.A.E.,  Mas, G., Figueroa, I., Borges, J. P. & Ávila, S. P. 2013. New records for the opisthobranch fauna of the Archipelago of the Azores (NE Atlantic Ocean). Marine Biodiversity Records, 6:e28.

62. Ohnheiser, L. T. & Malaquias, M. A. E. 2013. Systematic revision of the gastropod family Philinidae (Mollusca: Cephalaspidea) in the northeast Atlantic Ocean with emphasis on the Scandinavian peninsula. Zoological Journal of the Linnean Society, 167(2): 273-326.

61. Zabbey, N. & Malaquias, M. A. E. 2013. Epifauna diversity and ecology on intertidal flats in the tropical Niger Delta, with remarks on the gastropod species Haminoea orbignyanaJournal of the Marine Biological Association of the United Kingdom, 93(1): 249-257.   

60. Malaquias, M. A. E., Calado, G., Cruz, J. F. & Jensen, K. 2012. On the occurrence of the Caribbean sea slug Thuridilla mazda in the eastern Atlantic Ocean. Marine Biodiversity Records, Vol. 5, e50. doi:10.1017/S1755267211001023

59. Malaquias, M. A. E., Gosliner, T. M. & Cervera, J. L. 2011. Studies on opisthobranch molluscs: Proceedings of a symposium, World Congress of Malacology, Phuket, Thailand, July 2010. Introduction. Journal of Molluscan Studies, 77(4): 321.

58. Carmona, L., Malaquias, M. A. E., Gosliner, T. M., Pola, M. & Cervera, J. L. 2011. Amphi-Atlantic distributions and cryptic species in sacoglossan sea slugs.Journal of Molluscan Studies, 77(4): 401-412.

57. Pedro, N. C., Malaquias, M. A. E., Costa, A. C. & Ávila, S. P. 2011. Crimora papillata (Nudibranchia: Triophinae), a new record for the shallow marine molluscs of the Azores. Marine Biodiversity Records, 4: e37.

56. Zenetos, A., Katsanevakis, S., Poursanidis, D., Crocetta, F., Damalas, D., Apostolopoulos, G., Gravili, C., Vardala-Theodorou, E. & Malaquias, M. 2011. Marine alien species in Greek Seas: Additions and amendments by 2010.Mediterranean Marine Science, 12(1): 95-120.

55. Templado J. & Malaquias M.A.E. 2011. Familia Bullidae. Pp. 408-409. In: Gofas S., Moreno D. and Salas C. (coords.). Moluscos marinos de Andalucía. Málaga: Servicio de Publicaciones e Intercambio Cientifico, Universidad de Málaga.

54. Templado J., Malaquias M.A.E. & García García F. 2011. Familia Haminoeidae. Pp. 417-421. In: Gofas S., Moreno D. and Salas C. (coords.). Moluscos marinos de Andalucía. Málaga: Servicio de Publicaciones e Intercambio Cientifico, Universidad de Málaga.

53. Malaquias, M. A. E. 2011. Family Acteonidae d’Orbigny, 1842 – Review and commented. In: Severns, M.: Shells of the Hawaiian Islands – The Sea Shells: 390, pl. 178. ConchBooks, Hackenheim, Germany. 562p.

52. Malaquias, M. A. E. 2011. Family Bullinidae J. E. Gray, 1850 – Review and commented.In: Severns, M.: Shells of the Hawaiian Islands – The Sea Shells: 394, pl. 179. ConchBooks, Hackenheim, Germany. 562p.

51. Malaquias, M. A. E. 2011. Family Bullidae d’Orbigny, 1827 – Review and commented.In: Severns, M.: Shells of the Hawaiian Islands – The Sea Shells: 414, pl. 189. ConchBooks, Hackenheim, Germany. 562p.

50. Malaquias, M. A. E. 2011. Family Haminoeidae Pilsbry, 1895 – Review and commented.In: Severns, M.: Shells of the Hawaiian Islands – The Sea Shells: 414, pls 189–191. ConchBooks, Hackenheim, Germany. 562p.

49. Malaquias, M. A. E. 2011. Family Cylichnidae H. Adams & A. Adams, 1854 – Review and commented. In: Severns, M.: Shells of the Hawaiian Islands – The Sea Shells: 418, pl. 191. ConchBooks, Hackenheim, Germany. 562p.

48. Malaquias, M. A. E. 2010. Systematics, phylogeny, and natural history ofBullacta exarata (Philippi, 1849): an endemic cephalaspidean gastropod from the China Sea. Journal of Natural History, 44(33/34): 2015-2019.

47. Malaquias, M. A. E. 2009. Bermudas: no triângulo à procura de opistobrânquios [Bermuda: seeking opisthobranchs in the triangle]. Portugala, 14: 47-49.

46. Malaquias, M. A. E. & Reid, D. G. 2009. Tethyan vicariance, reliction and speciation: evidence from a global molecular phylogeny of the opisthobranch genus Bulla. Journal of Biogeography, 36: 1760-1777. 

45. Malaquias, M. A. E. 2009. Darwin: polvos, vinagreiras e fósseis na terra da morabeza [Darwin: octopus, sea hares, and fossils in the country of "morabeza"]. Portugala, 13: 3-4 [in Portuguese].

44. Ciavatta, M. L., Manzo, E., Nuzzo, G. Villani, G., Cimino, G., Cervera, J. L., Malaquias, M. A. E. & Gavagnin, M. 2009. Aplysiopsenes: an additional example of marine polyketides with a mixed acetate/propionate pathway. Tetrahedron Letters, 50: 527-529. 

43. Malaquias, M. A. E., Bericibar, E., & Reid, D. G. 2009. Reassessment of the trophic position of Bullidae (Gastropoda: Cephalaspidea) and the importance of diet in the evolution of cephalaspidean gastropods. Journal of Zoology, 277: 88-97. 

42. Malaquias, M. A. E., Dodds, J. M., Bouchet, P., & Reid, D. G. 2009. A molecular phylogeny of the Cephalaspidea sensu lato (Gastropoda: Euthyneura): Architectibranchia redefined and Runcinacea reinstated. Zoologica Scripta, 38(1): 23-41. 

41. Malaquias, M. A. E., Calado, G. P., Padula, V., Villani, G. & Cervera, J. L. 2009. Molluscan diversity in the North Atlantic Ocean: New records of opisthobranchs gastropods from the Archipelago of the Azores. Marine Biodiversity Records, 2: e38. 

40. Malaquias, M. A. E & Reid, D. G. 2008. Functional morphology of the gonoduct of the gastropod Bulla striata Bruguière, 1792 (Opisthobranchia: Cephalaspidea): evidence for a monaulic system. Acta Zoologica, 89: 205-210. 

39. Malaquias, M. A. E. & Reid, D. G. 2008. Systematic revision of the Recent species of Bullidae (Mollusca: Gastropoda: Cephalaspidea), with a molecular phylogenetic analysis. Zoological Journal of the Linnean Society, 153: 453–543. 

38. Malaquias, M. A. E. 2008. A short tale of a doctoral project: bubble-shells of the world. The Malacologist, 51: 24-29. 

37. Malaquias, M. A. E. 2008. A colecção malacológica do Museu Nacional de Ciências Naturais de Madrid  [The malacological collection of the Natural Sciences Museum of Madrid]. Portugala, 11: 5-6. [in Portuguese] 

36. Malaquias, M. A. E. 2007. Systematics, evolution and ecology of Bullidae (Mollusca, Gastropoda), with a molecular phylogeny of the order Cephalaspidea. PhD Thesis, University of London, January 2007, 276p. 

35. Malaquias, M. A. E. 2007. Gastropoda. In: Biodiversity in the fisheries of Algarve (south of Portugal). Borges, T.C. (ed.). Publisher: University of the Algarve, Faro: 196-241. 

34. Malaquias, M. A. E. 2007. Congresso Mundial de Malacologia 2007 – Antuérpia Bélgica [World Congress of Malacology – Antwerp, Belgium]. Portugala, 10: 9. [in Portuguese] 

33. Malaquias, M. A. E. 2006. 2º Workshop Internacional de Opistobrânquios [II International Workshop on Opisthobranchs]. Portugala, 8: 16. [in Portuguese] 

32. Malaquias, M. A. E. 2006. Flabellina ilidioi: uma espécie em homenagem a Ilídio Félix-Alves [Flabellina ilidioi: a species described in honour to Ilídio Félix-Alves]. Portugala, 7: 7. [in Portuguese] 

31. Malaquias, M. A. E. & Cervera, J. L. 2006. The genus Haminoea (Mollusca: Gastropoda: Cephalaspidea) in Portugal, with a review of the European species. Journal of Molluscan Studies, 72: 89-103. 

30. Malaquias, M. A. E., Bentes, L., Erzini, K. & Borges, T. C. 2006 Molluscan biodiversity caught by trawling fisheries: a case study in southern Portugal. Fisheries Management and Ecology, 13: 39-45. 

29. Coelho, R., Malaquias, M. A. E. & Calado, G. 2006. Calliopea bellula, a sacoglossan feeding upon egg masses of Haminoea orbignyana. Journal of the Marine Biological Association of the United Kingdom, 86: 423-424. 

28. Malaquias, M. A. E. & Sprung, M. 2005. Population biology of the cephalaspidean mollusc Haminoea orbygniana in a temperate coastal lagoon (Ria Formosa, Portugal). Estuarine Coastal and Shelf Science, 84: 767-772. 

27. Malaquias, M. A. E. 2005. As lesmas-do-mar nos Açores [Sea-slugs of the Azores]. Portugala, 5: 3-3. [in Portuguese] 

26. Malaquias, M. A. E. 2004. Congresso Mundial de Malacologia, Perth, Austrália, Julho de 2004 [World Congress of Malacology, Perth, Australia, July 2004]. Portugala, 4: 7-15. [in Portuguese] 

25. Malaquias, M. A. E., Condinho, S. Cervera, J. L. &. Sprung, M. 2004. Diet and feeding biology of Haminoea orbignyana (Férussac, 1828) (Mollusca: Gastropoda: Cephalaspidea). Journal of the Marine Biological Association of the United Kingdom, 84: 767-772. 

24. Malaquias, M. A. E. 2004. The opisthobranch molluscs described by the Reverend Robert Boog Watson from the Madeira Archipelago (Northeast Atlantic, Portugal). Journal of Conchology, 38: 231-240. 

23. Cervera, J. L., Calado, G., Gavaia, C., Malaquias, M. A. E., Templado, J., García-Gómez, J. C. & Megina, C. 2004. An annotated and updated checklist of the opisthobranchs (Mollusca: Gstropoda) from Spain and Portugal (including islands and archipelagos). Boletín Instituto Español de Oceanografía, 20: 1-122. 

22. Calado, G., Malaquias, M. A. E., Gavaia, C., Cervera, J. L., Megina, C., Dayrat, B., Camacho, Y., Pola, M. & Grande, C., 2004. New data on Opisthobranchs (Mollusca; Gastropoda) from the Southwestern coast of Portugal. Boletín Instituto Español de Oceanografía, 19: 199-204. 

21. Malaquias, M. A. E., Borges, J. P. & Borges, T. C. 2003. On the occurrence of Callumbonella suturalis (Philippi, 1836) in the South Atlantic coasts of Europe. Journal of Conchology, 38: 1-5. 

20. Ávila, S. P. and Malaquias, M. A. E. 2003. Biogeographycal relationships of the molluscan fauna of the Ormonde seamount (Gorringe Bank, Northeast Atlantic Ocean). Journal of Molluscan Studies, 69: 145-150. 

19. Gavaia, C., Malaquias, M. A. E., Calado, G. & Urgorri, V. 2003. New records of Portuguese opisthobranch molluscs. Journal of Conchology, 38: 101-118. 

18. Malaquias, M. A. E. 2003. Contribuição para a sistemática e ecologia do género Haminoea Turton and Kingston, 1830 (Mollusca: Opisthobranchia: Cephalaspidea) [Contribution to the systematics and ecology of the genus Haminoea Turton and Kingston, 1830 (Mollusca: Opisthobranchia: Cephalaspidea)]. MSc Thesis, University of Coimbra, Portugal, 121p. [in Portuguese] 

17. Malaquias, M. A. E., Calado, G. & Gavaia, C. 2003. Fauna Ibérica, moluscos opistobrânquios: Campanha de Sagres-Sines 2002 [Fauna Iberica, opisthobranchs molluscs: Campaign of Sagres-Sines 2002]. Portugala, 1: 11. [in Portuguese] 

16. Malaquias, M. A. E., Martínez, E. & Abreu, A. D. 2002. Cephalaspidea (Mollusca: Opisthobranchia) of the Madeira archipelago and Selvagens Islands. American Malacological Bulletin, 17: 65-83. 

15. Martínez, E, Malaquias, M. A. E & Cervera, L. 2002. Chelidonura africana Pruvot-Fol, 1953 (Mollusca, Gastropoda): proposed designation of a neotype. Journal of Conchology, 37: 349-353. 

14. Sprung, M. J., Afonso, C. M., Conduto, T., Machado, M., Malaquias, M. A. E.& Morenito, P. 2002. Evolução do macrobentos da Ria Formosa em zonas dragadas (EVODRAG). Relatório final [Succession of macrobenthic communities in recently dredged areas in Ria Formosa (EVODRAG). Final report]. Projecto IMAR/CCMAR – Universidade do Algarve / Instituto de Conservação da Natureza – Parque Natural da Ria Formosa, 22p+anexos. [in Portuguese] 

13. Malaquias, M. A. E. 2001. Updated and annotated checklist of the opisthobranch molluscs (excluding Thecosomata and Gymnosomata), from the Azores archipelago (North Atlantic Ocean, Portugal). Iberus, 19: 37-48. 

12. Malaquias, M. A. E., Cervera, J. L. Abreu, A. D. & López-González, P. 2001. The Opisthobranch Molluscs from Porto Santo Island (Madeira Archipelago, Northeastern Atlantic). Iberus, 19: 75-82. 

11. Borges, T. C., Costa, M. E., Erzini, K., Gonçalves, L., Malaquias, M., Olim, S., Pais, C., Ramos, J. & Sendão. J. C. 2001. Análise e avaliação do efeito da pesca de arrasto nas espécies demersais da costa do Algarve: quantificação e estudos bio-ecológicos. Relatório final [Study of the impact of trawling fisheries in the Algarve: evaluation and bio-ecological survey. Final report], Fundação para a Ciência e Tecnologia, Contrato No.21/REGII/96, 288p. [in Portuguse] 

10. Malaquias, M. A. E. 2000. Additions to the knowledge of the opisthobranch molluscs of Selvagens Islands, NE Atlantic, Portugal. Arquipélago, Life and Marine Sciences, Supplement 2(A): 89-97.

9.  Malaquias, M. A. E. & Morenito, P. M. 2000. The opisthobranch (Mollusca: Gastropoda) of a coastal lagoon Ria Formosa in southern Portugal. Bolletino Malacologico, 36: 117-124. 

8. Sprung, M. J., Machado, M., Malaquias, M. A. E.& Morenito, P. 2000. Impacte das dragagens nas comunidades bentónicas da Ria Formosa. Relatório final [Impact of dredging activities in the benthic communities of Ria Formosa. Final report]. Projecto IMAR/CCMAR – Universidade do Algarve / Instituto de Conservação da Natureza – Parque Natural da Ria Formosa, 50p. [in Portuguse] 

7. Malaquias, M. A. E. & Calado, G. 1997. The malacological fauna of Selvagens Archipelago. 1 Opisthobranch Molluscs. Boletim do Museu Municipal do Funchal, 29: 149-170. 

6. Malaquias, M. A. E. 1996. Opistobrânquios (Mollusca Gastropoda) do arquipélago da Madeira e das ilhas Selvagens [Opisthobranchs (Mollusca: Gastropoda) of Madeira archipelago and Selvagens Islands]. BSc Thesis, University of the Algarve, Portugal, 76p. [in Portuguese] 

5. Abreu, A. D., Araújo, R, Biscoito, M. J. e Malaquias, M. A. E. 1995. Fauna Marinha da Madeira [Marine fauna and flora of Madeira archipelago]. Jornal da Madeira (Suplemento). 108p. [in Portuguese] 

4. Malaquias, M. A. E. 1991. Ria Formosa – Artes e Vidas [Ria Formosa – Life and arts]. HALIA, Boletim da Sociedade Portuguesa de Malacologia, 5-III Ser.: 2-6. [in Portuguese] 

3. Malaquias, M. A. E. 1991. Ria Formosa – Localização e características [Ria Formosa – Location and features]. HALIA, Boletim da Sociedade Portuguesa de Malacologia, 4-III Ser.: 16-18. [in Portuguese] 

2. Malaquias, M. A. E. 1989. Contribuição para o inventário da fauna malacológica da linha de Cascais (Continuação) [Contribution to the knowledge of the malacological fauna of Cascais’ shores (continuation)]. HALIA, Boletim da Sociedade Portuguesa de Malacologia, 24-NS: 12-12. [in Portuguese] 

1. Malaquias, M. A. E. 1989. Contribuição para o inventário da fauna malacológica da linha de Cascais [Contribution to the knowledge of the malacological fauna of Cascais’ shores]. HALIA, Boletim da Sociedade Portuguesa de Malacologia, 23-NS: 7-13. [in Portuguese]

 

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3. Diversity, systematics and speciation of the genus Haminoea in the Atlantic and Eastern Pacific Oceans

4. Phylogeography of the gastropod Bulla occidentalis in the tropical Western Atlantic Ocean

5. Diversity, systematics and trophic biology of the deep sea gastropod genus Scaphander

6. Systematics and phylogeny of Cephalaspidea gastropods

7. Sea slugs diversity (nudibranchs & others) of Norway

8. Biogeography and evolution of the Indian Ocean biota

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 2007: PhD in systematics and evolution; Queen Mary College, University of London and Natural History Museum, London (thesis title: Systematics, evolution and ecology of Bullidae (Mollusca, Gastropoda), with a molecular phylogeny of the order Cephalaspidea).

2003: MSc in Ecology; University of Coimbra, Portugal (thesis title: Systematics and ecology of the genus Haminoea Turton and Kingston, 1830 (Mollusca: Opisthobranchia: Cephalaspidea)). 

1996: BSc (“Licenciatura”) in Marine Biology; University of the Algarve, Portugal (thesis title: Opisthobranchs (Mollusca: Gastropoda) of the Madeira archipelago and Selvagens Islands).