• E-mailSteffen.Roth@uib.no
  • Phone+47 55 58 29 11+47 975 51 389
  • Visitor Address
    Allégaten 41
    5007 Bergen
  • Postal Address
    Postboks 7800
    5020 Bergen

- Management of the insect collection

- Phylogeny, Ecology, and Faunistics of Heteroptera

 - Amphibiens in Norway

My homepage


Academic article
  • Show author(s) (2023). The evolution of female-biased genital diversity in bedbugs (Cimicidae). Evolution. 329-341.
  • Show author(s) (2023). Primicimex cavernis Barber 1941 (Heteroptera: Cimicidae): A rare parasite on a very abundant host. Insect Conservation and Diversity. 163-167.
  • Show author(s) (2023). First records of bat-associated Cimex lectularius (Cimicidae, Heteroptera) for Armenia and Georgia. Caucasiana.
  • Show author(s) (2023). Contribution to the Cimicidae (Hemiptera: Heteroptera) fauna from Argentina: biology and geographical distribution. Zootaxa. 587-594.
  • Show author(s) (2021). Type of broadleaf forest matters most for ptyctimous mite communities (Acari, Oribatida) in Norway. Biodiversity and Conservation. 2929-2953.
  • Show author(s) (2021). The Arthropod Fauna of Oak (Quercus spp., Fagaceae) Canopies in Norway. Diversity. 31 pages.
  • Show author(s) (2021). High Diversity of Mites (Acari: Oribatida, Mesostigmata) Supports the High Conservation Value of a Broadleaf Forest in Eastern Norway. Forests.
  • Show author(s) (2021). A forest pool as a habitat island for mites in a limestone forest in Southern Norway. Diversity. 578.
  • Show author(s) (2020). The firebug Pyrrhocoris apterus (Linnaeus, 1758) (Hemiptera, Heteroptera) new to the Norwegian fauna – with an explosive expansion in Northern Europe. Norwegian Journal of Entomology. 81-90.
  • Show author(s) (2019). Diverse mite communities (Acari: Oribatida, Mesostigmata) from a broadleaf forest in western Norway. Annales Zoologici Fennici. 121-136.
  • Show author(s) (2019). Bedbugs (Cimicidae) evolved before their bat hosts and did not co-speciate with ancient humans. Current Biology. 1847-1853.
  • Show author(s) (2018). Wanzenfunde (Insecta: Heteroptera) aus Nordhessen anlässlich des 43. Treffens der „Arbeitsgruppe Mitteleuropäischer Heteropterologen“ im August 2017. Philippia. 219-264.
  • Show author(s) (2018). Genetic diversity of common toads (Bufo bufo) along the norwegian coast: Disjunct distribution of locally dominant haplotypes. Herpetological Journal. 127-133.
  • Show author(s) (2018). Co-speciation in bedbug Wolbachia parallel the pattern in nematode hosts. Scientific Reports. 1-9.
  • Show author(s) (2018). Cimex pipistrelli complex new to Norway and additional records of bed bug species (Cimicidae, Heteroptera) from Norway and Sweden. Norwegian Journal of Entomology. 118-126.
  • Show author(s) (2016). Large populations of the common toad Bufo bufo in Norway. Herpetology Notes. 325-330.
  • Show author(s) (2016). High genetic diversity of common toad (Bufo bufo) populations under strong natural fragmentation on a Northern archipelago. Ecology and Evolution. 1626-1636.
  • Show author(s) (2015). Molecular evidence places the swallow bug genus Oeciacus Stål within the bat and bed bug genus Cimex Linnaeus (Heteroptera: Cimicidae). Systematic Entomology. 652-665.
  • Show author(s) (2014). Fieberocapsus flaveolus (Reuter,1870)(Miridae, Heteroptera) - ein bemerkenswerter Wanzenfund für Sachsen-Anhalt. Entomologische Mitteilungen Sachsen-Anhalt. 47-48.
  • Show author(s) (2014). Biodiversity, ecology, and behavior of the recently discovered insect order Mantophasmatodea . Frontiers in Zoology.
  • Show author(s) (2014). A survey of aquatic and terrestrial Heteroptera in northern Europe with special regard to Finnmark, Norway (and adjacent regions). Norwegian Journal of Entomology. 99-116.
  • Show author(s) (2013). Spatiotemporal analysis of predation by carabid beetles (Carabidae) on nematode infected and uninfected slugs in the field. PLOS ONE. 14 pages.
  • Show author(s) (2013). Predation by carabid beetles on the invasive slug Arion vulgaris in an agricultural semi-field experiment. Bulletin of entomological research. 225-232.
  • Show author(s) (2013). Distribution and spread of the invasive slug Arion vulgaris Moquin - Tandon in Norway. Fauna Norvegica. 13-26.
  • Show author(s) (2012). Some notes on reproductive biology and mating behaviour of Arion vulgaris Moquin-Tandon 1855 in Norway including a mating experiment with a hybrid of Arion rufus (Linnaeus 1758) x ater (Linnaeus 1758). Journal of Conchology. 249-257.
  • Show author(s) (2012). Peptidomics-based phylogeny and biogeography of mantophasmatodea (Hexapoda). Systematic Biology. 609-629.
  • Show author(s) (2012). Insects inhabiting freshwater and humid habitats in Finnmark, northern Norway. Norwegian Journal of Entomology. 91-107.
  • Show author(s) (2012). Ectoparasite loads of the Central American Whiptail Lizard Ameiva festiva (Squamata: Teiidae). Herpetological Journal. 151-155.
  • Show author(s) (2012). A „winter“ inscet- the third record of Pachycerina pulchra (Loew, 1850) (Diptera, Brachycera, Fam. Lauxaniidae) from Norway. Dipterists Digest. 184.
  • Show author(s) (2009). Sexual dimorphism in winter survival rate differs little between damselbug species (Heteroptera: Nabidae). European Journal of Entomology. 37-41.
  • Show author(s) (2009). Die Wanzen Norwegens: Arten, Erfassungsstand und Rote Liste. Heteropteron. 7-13.
  • Show author(s) (2009). A proteomic approach for studying insect phylogeny: CAPA peptides of ancient insect taxa (Dictyoptera, Blattoptera) as a test case. BMC Evolutionary Biology.
  • Show author(s) (2008). Odour-mediated sexual attraction in nabids (Heteroptera : Nabidae). European Journal of Entomology. 159-162.
Popular scientific lecture
  • Show author(s) (2013). Biologi og bekjemplse av Brunskogsnegl (Arion vulgaris).
Academic lecture
  • Show author(s) (2018). The evolution of specialisation and generalism in bedbugs (Cimicidae) .
  • Show author(s) (2014). A molecular phylogeny of the bed bug family (Cimicidae): preliminary results.
  • Show author(s) (2012). Kartlegging av insekter knyttet til vann og fuktige habitater i Finnmark.
  • Show author(s) (2009). Die Wanzen Norwegens:Arten, Rote Liste und Erfassungsstand (Heteroptera in Norway: species, Red Data List and knowledge).
Popular scientific article
  • Show author(s) (2012). Vannelskende insekter i Finnmark. Årbok for Universitetsmuseet i Bergen. 92-103.
  • Show author(s) (2012). Neuer Wein aus alten Schläuchen: DNA-Untersuchungen an Sammlungsmaterial. Heteropteron. 5-9.
  • Show author(s) (2012). Buchbesprechung "Suomen Luteet" (Die Wanzen Finnlands). Heteropteron. 2-2.
  • Show author(s) (2009). Mystisk insekt.
  • Show author(s) (2009). Fyrste Funn i fylket.
Museum exhibition
  • Show author(s) (2019). Insekter - en million suksesshistorier.
  • Show author(s) (2019). De første virveldyr på land.
  • Show author(s) (2009). The genetic structure of common toad (Bufo bufo) populations under long-term, natural fragmentation.

More information in national current research information system (CRIStin)


  • Phylogeny and ecology of the Cimicidae: Bedbugs, or Cimicidae, copulate in a spectacular way: males pierce the cuticle of the female and transfer their sperm into the female body (traumatic insemination). In at least two cimicid species this reproductive mode has been shown or implicated to result in sexual conflict, a process that is known to accelerate the rate of evolutionary change. The aims of this study are a) revealing the molecular phylogenetic relationship within the Cimicidae and b) to reconstructing the evolution of sexual organs, particularly the female copulatory organ which seems to ameliorate the damage caused by the male.

  • Evolution of reproductive systems in damselbugs (Nabidae) - Linking behaviour, morphology and phylogeny
  • Phylogeny and ecology of Mantophasmatodea
  • Faunistics of Heteroptera in Norway
  • Population dynamics of Amphibians in Norway

My homepage