Sigurd Stefansson
- E-mailSigurd.Stefansson@uib.no
- Phone+47 55 58 44 70+47 930 84 646
- Visitor AddressThormøhlens gate 53 A/B5006 Bergen
- Postal AddressPostboks 78035020 Bergen
Professor Sigurd O. Stefansson (December, 2017 ISI statistic 158, H-index 43) has long-term research experience in fish growth and development, reproduction and adaptation to the environment, both in salmonids and marine species, using a wide range of tools. Central in Stefansson’s research is the process of salmon smoltification, and his research includes both basic research and collaboration with the aquaculture industry. Research topics include the environmental, genetic and endocrine control of smoltification and basic mechanisms of adaptation to seawater.
Professional experience
Institute of Marine Research (IMR), Matre Aquaculture Station, Research fellow, Norwegian Fisheries Research Council 'Miljøforholdene i settefiskproduksjon', 07.1986 - 08.1988
Dept. of Fisheries Biology, UiB, Associate Professor (Amanuensis), 08.1988 - 06.1990
IMR, Matre Aquaculture Station, Senior Scientists, 01.07. - 31.12.1990
Dept. of Fisheries and Marine Biology, UiB, Associate professor 12.1990 – 03.2001
Department of Fisheries and Oceans, Biological Station, St. Andrews, Canada, post-doc. (Norwegian Fisheries Research Council), 05. - 12.1992
Dept. of Biology, UiB, Professor, 01.04.2001 – present
Univ. of Massachusetts at Amherst and Conte Anadromous Fish Research Center, Turners Falls, MA, USA, Sabbatical, 08.2003 – 07.2004
Completed Univ. of Bergen programme on teaching and pedagocics, 1994
Member of Dept. of Biology programme board for teaching (several appointments)
Chair of Education, Dept. of Biology, 2016
Chair of WP1 bioCEED, 2016
Head of the study programme “Havbruk og Sjømat” (Aquaculture and Seafood), 2016 – ongoing (integrated master/siv. ing.)
Courses
BIF 100 Innføring i fiskehelse og havbruk
Course responsible, teaching, assessment, quality assurance
BIF 101 Organismal Biology in Aquatic Medicine and Aquaculture
Course responsible, teaching, assessment, quality assurance
BIO 104 Comparative physiology
Course responsible, teaching, assessment, quality assurance
BIO 203 Introduction to Aquaculture
Teaching, assessment, quality assurance
BIO 291 Fish Biology II - Fish Physiology
Teaching, assessment, quality assurance
BIO 304 Specialisation in Aquaculture Biology
Teaching, assessment, quality assurance
SupervisionCand. scient. and Master students:
43 students completed their degree, 3 currently under supervision
Dr. scient and PhD students:
12 students completed their degree, 3 students currently under supervision
Post-doc researchers:
3 finished
- (2022). Temperature and size-dependency of lumpfish (Cyclopterus lumpus) oxygen requirement and tolerance. Aquaculture. 10 pages.
- (2022). Lumpfish Cyclopterus lumpus reproduction: Pituitary gene expression, physiological and morphological changes accompanying gonadal maturation. Aquaculture. 1-15.
- (2022). Investigation of growth performance of post-smolt Atlantic salmon (Salmo salar L.) in semi closed containment system: A big-scale benchmark study. Aquaculture Research. 4178-4189.
- (2022). Increased Thermal Challenges Differentially Modulate Neural Plasticity and Stress Responses in Post-Smolt Atlantic Salmon (Salmo salar). Frontiers in Marine Science. 16 pages.
- (2022). Exposure to cold temperatures differentially modulates neural plasticity and stress responses in post-smolt Atlantic salmon (Salmo salar). Aquaculture. 11 pages.
- (2020). Performance and welfare of Atlantic salmon, Salmo salar L. post-smolts in recirculating aquaculture systems: Importance of salinity and water velocity. Journal of the World Aquaculture Society. 373-392.
- (2019). The use of photoperiods to provide year round spawning in lumpfish Cyclopterus lumpus. Comparative Biochemistry and Physiology A. 62-70.
- (2018). The effect of continuous light and compressed photoperiods on growth and maturation in lumpfish Cyclopterus lumpus. Aquaculture. 166-172.
- (2018). High fish density delays wound healing in Atlantic salmon (Salmo salar). Scientific Reports. 13 pages.
- (2018). Atlantic salmon post-smolts adapted for a longer time to seawater develop an effective humoral and cellular immune response against Salmonid alphavirus. Fish and Shellfish Immunology. 579-590.
- (2018). Atlantic salmon adapted to seawater for 9 weeks develop a robust immune response to salmonid alphavirus upon bath challenge. Fish and Shellfish Immunology. 573-583.
- (2017). Triploid atlantic salmon (Salmo salar L.) postsmolts accumulate prevalence more slowly than diploid salmon following bath challenge with salmonid alphavirus subtype 3. PLOS ONE. 1-12.
- (2017). The effect of continuous light at low temperatures on growth in Atlantic salmon reared in commercial size sea pens. Aquaculture. 645-651.
- (2017). Stocking density limits for post-smolt Atlantic salmon (Salmo salar L.) emphasis on production performance and welfare. Aquaculture. 363-370.
- (2017). Immune gene profiles in Atlantic salmon (salmo salar L.) post-smolts infected with SAV3 by bath-challenge show a delayed response and lower levels of gene transcription compared to injected fish. Fish and Shellfish Immunology. 320-331.
- (2016). The effect of low temperatures and photoperiods on growth and vertebra morphometry in Atlantic salmon. Aquaculture International. 1421-1434.
- (2016). Relationship between viral dose and outcome of infection in Atlantic salmon, Salmo salar L., post-smolts bath-challenged with salmonid alphavirus subtype 3. Veterinary research (VR). 1-13.
- (2016). Impact of fish density and specific water flow on skin properties in Atlantic salmon (Salmo salar L.) post-smolts. Aquaculture. 629-637.
- (2016). Identification of a novel leptin receptor duplicate in Atlantic salmon: Expression analyses in different life stages and in response to feeding status. General and Comparative Endocrinology. 108-119.
- (2016). Growth and Smoltification of Three Norwegian Strains of Atlantic Salmon Salmo salar Reared under Different Thermal Regimes. Rivers Research and Applications: an international journal devoted to river research and management. 1800-1807.
- (2016). Effects of temperature on the final stages of sexual maturation in Atlantic salmon (Salmo salar L.). Fish Physiology & Biochemistry. 895-907.
- (2016). Atlantic salmon (Salmo salar L.) post-smolts challenged two or nine weeks after seawater-transfer show differences in their susceptibility to salmonid alphavirus subtype 3 (SAV3). Virology Journal. 14 pages.
- (2015). Roles of leptin and ghrelin in adipogenesis and lipid metabolism of rainbow trout adipocytes in vitro. Comparative Biochemistry and Physiology A. 40-48.
- (2014). Plasma growth hormone-binding protein levels in Atlantic salmon Salmo salar during smoltification and seawater transfer. Journal of Fish Biology. 1279-1296.
- (2014). Photoperiod and temperature effects on growth and maturation of pre- and post-smolt Atlantic salmon. Aquaculture International. 1331-1345.
- (2014). Osmoregulation in Atlantic salmon Salmo salar smolts transferred to seawater at different temperatures. Journal of Fish Biology. 1163-1176.
- (2014). Osmoregulation and growth in offspring of wild Atlantic salmon at different temperatures. Environmental Biology of Fishes. 285-296.
- (2014). Development of intestinal ion-transporting mechanisms during smoltification and seawater acclimation in Atlantic salmon Salmo salar. Journal of Fish Biology. 1227-1252.
- (2013). Seawater tolerance and post-smolt migration of wild Atlantic salmon Salmo salar × brown trout S. trutta hybrid smolts. Journal of Fish Biology. 206-227.
- (2013). Molecular cloning and genomic characterization of novel Leptin-like genes in salmonids provide new insight into the evolution of the Leptin gene family. General and Comparative Endocrinology. 48-59.
- (2013). Low light intensity can reduce Atlantic salmon smolt quality. Aquaculture. 19-24.
- (2013). Long-term effects of photoperiod, temperature and their interaction on growth, gill Na+, K+-ATPase activity, seawater tolerance and plasma growth-hormone levels in Atlantic salmon Salmo salar. Journal of Fish Biology. 1197-1209.
- (2013). Aluminum exposure impacts brain plasticity and behavior in Atlantic salmon (Salmo salar). Journal of Experimental Biology. 3148-3155.
- (2012). The role of growth hormone in growth, lipid homeostasis, energy utilization and partitioning in rainbow trout: Interactions with leptin, ghrelin and insulin-like growth factor I. General and Comparative Endocrinology. 153-162.
- (2012). Salmon lice-induced mortality of Atlantic salmon postsmolts experiencing episodic acidification and recovery in freshwater. Aquaculture. 193-199.
- (2012). Recovery of Atlantic salmon smolts following aluminum exposure defined by changes in blood physiology and seawater tolerance. Aquaculture. 232-240.
- (2012). Larval rearing environment influences the physiological adaptation in juvenile Atlantic cod, Gadus morhua. Aquaculture International. 467-479.
- (2012). Isolation and characterization of melanopsin photoreceptors of atlantic salmon (Salmo salar). Journal of Comparative Neurology. 3727-3744.
- (2012). Growth, osmoregulation and endocrine changes in wild Atlantic salmon smolts and post-smolts during marine migration. Aquaculture. 127-136.
- (2011). Leptin reduces Atlantic salmon growth through the central pro-opiomelanocortin pathway. Comparative Biochemistry and Physiology A. 79-86.
- (2011). Involvement of Prop1 homeobox gene in the early development of fish pituitary gland. General and Comparative Endocrinology. 332-340.
- (2011). Environmental endocrinology of salmon smoltification. General and Comparative Endocrinology. 290-298.
- (2011). Effect of rearing temperatures on the growth and maturation of Arctic charr (Salvelinus alpinus) during juvenile and on-growing periods. Aquaculture Research. 221-229.
- (2011). Corticotropin-Releasing Factor Neurogenesis during Midlife Development in Salmon: Genetic, Environmental and Thyroid Hormone Regulation. Journal of neuroendocrinology. 733-741.
- (2011). A retrospective approach to fractionize variation in body mass of Atlantic cod Gadus morhua. Journal of Fish Biology. 251-264.
- (2010). The influence of first-feeding diet on the Atlantic cod Gadus morhua phenotype: survival, development and long-term consequences for growth. Journal of Fish Biology. 1-19.
- (2010). Ontogenetic effects of diet during early development on growth performance, myosin mRNA expression and metabolic enzyme activity in Atlantic cod juveniles reared at different salinities. Comparative Biochemistry and Physiology A. 102-109.
- (2010). Leptin and leptin receptor genes in Atlantic salmon: Cloning, phylogeny, tissue distribution and expression correlated to long-term feeding status. General and Comparative Endocrinology. 55-70.
- (2010). Effects of acidic water and aluminum exposure on gill Na+, K+-ATPase alpha-subunit isoforms, enzyme activity, physiology and return rates in Atlantic salmon (Salmo salar L.). Aquatic Toxicology. 250-259.
- (2009). Long-term effect of photoperiod manipulation on growth, maturation and flesh quality in Atlantic halibut. Aquaculture Research. 1260-1269.
- (2009). Influence of sea temperature and initial marine feeding on survival of Atlantic salmon Salmo salar post-smolts from the Rivers Orkla and Hals, Norway. Journal of Fish Biology. 1532-1548.
- (2009). Food-deprivation, compensatory growth and hydro-mineral balance in Atlantic salmon (Salmo salar) post-smolts in sea water. Aquaculture. 243-249.
- (2009). Effects of fish density on growth and smoltification in intensive production of Atlantic salmon (Salmo salar L.). Aquaculture. 236-241.
- (2009). Effects of chronic and periodic exposure to ammonia on growth and blood physiology in juvenile turbot (Scophthalmus maximus). Aquaculture. 45-50.
- (2009). Compensatory growth in Atlantic halibut: Effect of starvation and subsequent feeding on growth, maturation, feed utilization and flesh quality. Aquaculture. 304-310.
- (2009). Characterization, tissue distribution, and regulation of agouti-related protein (AgRP), cocaine- and amphetamine-regulated transcript (CART) and neuropeptide Y (NPY) in Atlantic salmon (Salmo salar). General and Comparative Endocrinology. 160-171.
- (2009). A homologous salmonid leptin radioimmunoassay indicates elevated plasma leptin levels during fasting of rainbow trout. General and Comparative Endocrinology. 307-312.
- (2008). The effect of temperature and fish size on growth, feed intake, food conversion efficiency and stomach evacuation rate of Atlantic salmon post-smolts. Aquaculture. 36-42.
- (2008). The combined effects of temperature and GnRHa treatment on the final stages of sexual maturation in Atlantic salmon (Salmo salar L.) females. Fish Physiology & Biochemistry. 289-298.
- (2008). Plasma insulin-like growth factor-I concentrations and growth in juvenile halibut (Hippoglossus hippoglossus): Effects of photoperiods and feeding regimes. Comparative Biochemistry and Physiology A. 66-70.
- (2008). Nutritional status of Atlantic salmon (Salmo salar) broodstock. Maternal effects on early expression of leptin, IGF-I, IGF-IR, and GHR. Cybium. 62-63.
- (2008). Interactive effects of ammonia and oxygen on growth and physiological status of juvenile Atlantic cod (Gadus morhua). Aquaculture. 292-299.
- (2008). Independent and dynamic reallocation of pitx gene expression during vertebrate evolution, with emphasis on fish pituitary development. Gene. 19-26.
- (2008). Endocrine systems in juvenile anadromous and landlocked Atlantic salmon (Salmo salar): Seasonal development and seawater acclimation. General and Comparative Endocrinology. 762-772.
- (2008). Effect of ambient oxygen on growth and reproduction in Nile tilapia (Oreochromis niloticus). Canadian Journal of Fisheries and Aquatic Sciences. 1413-1424.
- (2008). Daily endocrine profiles in parr and smolt Atlantic salmon. Comparative Biochemistry and Physiology A. 698-704.
- (2008). Branchial expression patterns of claudin isoforms in Atlantic salmon during seawater acclimation and smoltification. American Journal of Physiology. Regulatory Integrative and Comparative Physiology. R1563-R1574.
- (2007). Temperature and salinity effects on plasma insulin-like growth factor-I concentrations and growth in juvenile turbot (Scophthalmus maximus). Aquaculture. 546-552.
- (2007). Salmon lice or suboptimal water quality - Reasons for reduced postsmolt survival? Aquaculture. 374-383.
- (2007). Persistent growth effects of temperature and photoperiod in Atlantic cod Gadus morhua. Journal of Fish Biology. 1371-1382.
- (2007). New haemoglobin genotypes in Atlantic cod, Gadus morhua: Possible relation with growth. Comparative Biochemistry and Physiology A. 955-960.
- (2007). Molecular mechanisms of contiuous light inhibition of Atlantic salmon parr-smolt transformation. Aquaculture. 235-245.
- (2007). Interactive effects of oxygen saturation blood physiology and ammonia on growth and in juvenile turbot. Aquaculture. 244-251.
- (2007). Interaction between temperature and photoperiod on growth and feeding of Atlantic cod (Gadus morhua): possible secondary effects. Canadian Journal of Fisheries and Aquatic Sciences. 239-248.
- (2007). Impacts of elevated water carbon dioxide partial pressure at two temperatures on Atlantic salmon (Salmo salar L.) parr growth and haematology. Aquaculture. 241-249.
- (2007). Exposure to moderate acid water and aluminum reduces Atlantic salmon post-smolt survival. Aquaculture. 360-373.
- (2007). Exposure to continuous light disrupts retinal innervation of the preoptic nucleus during parr-smolt transformation in Atlantic salmon. Aquaculture. 345-349.
- (2007). Differential expression of gill Na+,K+-ATPase alpha- and beta-subunits, Na+,K+,2Cl(-) cotransporter and CFTR anion channel in juvenile anadromous and landlocked Atlantic salmon Salmo salar. Journal of Experimental Biology.
- (2006). The interrelation between temperature regimes and fish size in juvenilie Atlantic cod (Gadus morhua). effects on growth and feed conversion efficiency. Fish Physiology & Biochemistry. 347-361.
- (2006). Short- and long-term differences in growth, feed conversion efficiency and deformities in juvenile Atlantic cod (Gadus morhua) startfed on rotifers or zooplankton. Aquaculture Research. 1015-1027.
- (2006). Growth, feed conversion efficiency and growth heterogeneity in Atlantic halibut (Hippoglossus hippoglossus) reared at three different photoperiods. Aquaculture Research. 1099-1106.
- (2006). Effects of dietary soybeanmeal and photoperiod cycle on osmoregulation following seawater exposure in Atlantic salmon smolts. Journal of Fish Biology. 1396-1426.
- (2006). Does malpigmentation improve growth in juvenile turbot (Scophthalmus maximus Rafinesque) and halibut (Hippoglossus hippoglossus L.)? Aquaculture Research. 306-312.
- (2006). Compensatory growth and fillet crude composition in juvenile Atlantic halibut: Effects of short term starvation periods and subsequent feeding. Aquaculture. 109-117.
- (2005). The interrelation between temperature regimes and fish size in juvenile Atlantic cod (Gadus morhua): effects on growth and feed conversion efficiency. Fish Physiology & Biochemistry. 347-361.
- (2005). Genotypic response to photoperiod treatment in Atlantic cod (Gadus morhua). Aquaculture. 525-532.
- (2004). Influence of exogenous thyroxine on plasma melatonin in juvenile Atlantic salmon (Salmo salar). Comparative Biochemistry and Physiology - Part B: Biochemistry & Molecular Biology. 43-47.
- (2004). Effects of 3-thia fatty acids on feed intake,growth, tissue fatty acid composition, beta-oxidation and Na,K-ATPase activity in Atlantic salmon. ?. 657-668.
- (2003). Seawater adaptation and growth of post-smolt Atlantic salmon (Salmo salar) of wild and farmed strains. Aquaculture. 367-384.
- (2003). Osmoregulation, feed intake, growth and growth hormone levels in 0+ Atlantic salmon (Salmo salar L.) transferred to seawater at different stages of smolt development. Aquaculture. 167-187.
- (2001). Variation in food intake, food conversion efficiency and growth of juvenile turbot from different geographic strains. Journal of Fish Biology. 449-454.
- (2001). The interaction of temperature and salinity on growth and food conversion in juvenile turbot (Scophthalmus maximus). Aquaculture. 353-367.
- (2001). Selection or adaptation: Differences in growth performance of juvenile turbot (Scophthalmus maximus Rafinesque) from two close-by localities off Norway. Sarsia. 43-51.
- (2001). Selection or adaptation: Differences in growth performance of juvenile turbot (Scophthalmus maximus Rafinesque) from two close-by localities off Norway. Sarsia. 43-51.
- (2001). Daily profiles of plasma free thyroxine, total thyroxine, total triiodothyronine, and growth hormone in Atlantic salmon parr and smolts. Fish Physiology & Biochemistry.
- (2001). Comparisons of RNA/DNA ratios, growth and metabolism of different populations of juvenile turbot Scophthalmus maximus reared at four temperatures. Journal of the World Aquaculture Society. 1-10.
- (2000). The effect of different initial size distribution on the growth of Atlantic halibut (Hippoglossus hippoglossus L.). Journal of Fish Biology. 826-836.
- (2000). RNA/DNA ratios, growth and metabolism of different populations of juvenile turbot Scophthalmus maximus reared at four temperatures. Journal of the World Aquaculture Society.
- (2000). Intraspecific differences in physiological efficiency of juvenile Atlantic halibut Hippoglossus hippoglossus L. Journal of the World Aquaculture Society. 285-296.
- (2000). Interaction of temperature and photoperiod on growth of Atlantic halibut Hippoglossus hippoglossus L. Aquaculture Research. 219-227.
- (2000). Hemoglobin genotypes of turbot (Schophthalmus maximus): consequences for growth and variations in optimal temperature for growth. Fish Physiology & Biochemistry. 75-81.
- (2000). Growth of juvenile halibut (Hippoglossus hippoglossus) under different photoperiods. Aquaculture. 119-128.
- (2000). Geographic variation in growth and food conversion efficiency of juvenile Atlantic halibut related to latitude. Journal of Fish Biology. 279-294.
- (2000). Free plasma thyroxine levels in coho salmon, Oncorhynchus kisutch, during parr-smolt transformation: comparison with total thyroxine, total triiodothyronine and growth hormone levels. Fish Physiology & Biochemistry. 45-50.
- (2000). Environmental influences on melatonin secretion in Atlantic cod (Gadus morhua L.) and their relevance to commercial culture. Fish Physiology & Biochemistry. 191-200.
- (2000). Effects of continuous additional light on growth and sexual maturity in Atlantic salmon, Salmo salar, reared in sea cages. Aquaculture. 337-349.
- (2000). Countergradient variation in growth and growth efficiency of juvenile turbot. Journal of Fish Biology. 1213-1226.
- (1999). Temperature- and size-dependent growth of larval and early juvenile Atlantic cod (Gadus morhua): a comparative study of Norwegian coastal cod and northeast Arctic cod. Canadian Journal of Fisheries and Aquatic Sciences. 2099-2111.
- (1998). Effects of size grading on growth and survival of juvenile turbot at two temperatures. Aquaculture International. 19-32.
- (1998). Abrupt changes in photoperiod affect age at maturity, timing of ovulation and plasma testosterone and oestradiol-17 profiles in Atlantic salmon, Salmo salar. Aquaculture. 85-98.
- (1997). Effects of exposure to extended photoperiods during the first winter on long-term growth and age at first maturity in turbot (Scophthalmus macimus). Aquaculture. 125-141.
- (1995). Photoperiod Regulation of Plasma Growth Hormone Levels during Parr-Smolt Transformation of Atlantic Salmon: Implications for Hypoosmoregulatiory Ability and Growth. General and Comparative Endocrinology. 73-82.
- (1995). Growth, oxygen consumption and activity of juvenile turbot (Scophthalamus maximus) reared under different temperatures and photoperiods. Netherlands Journal of Sea Research. 149-159.
- (1995). Growth of juvenile halibut (Hippoglossus hippoglossus L.) related to temperature, day length and feeding regime. Netherlands Journal of Sea Research. 139-147.
- (1995). Development of salinity tolerance in underyearling smolts of Atlantic salmon (Salmo salar) reared under different photoperiods. Canadian Journal of Fisheries and Aquatic Sciences. 243-251.
- (1995). Development of salinity tolerance in underyearling smolts of Atlantic salmon (Salmo salar) reared under different photoperiods. Canadian Journal of Fisheries and Aquatic Sciences. 243-251.
- (1995). Changes in serum protein and IgM concentration during smolting and early post-smolt period in vaccinated and unvaccinated Atlantic salmon (Salmo salar L.). Fish and Shellfish Immunology. 211-221.
- (1994). The interrelation between photo-period, growth hormone, and sexual maturation of adult Atlantic salmon (Salmo salar). General and Comparative Endocrinology. 70-81.
- (1994). Predation and dietary habits as possible explanations for poor return rates of stocked seatrout from the river Aurland. Norsk Geografisk Tidsskrift. 51-53.
- (1994). Effects of photoperiod and temperature on growth and parr-smolt transformation in Atlantic salmon (Salmo salar L.) and subsequent performance in seawater. Aquaculture. 13-27.
- (1994). Effect of reduced daylength on growth, sexual maturation and smoltification in Atlantic salmon (Salmo salar) underyearlings. Aquaculture. 289-230.
- (1994). Effect of a simulated winter of different duration on growth and smoltification in Atlantic salmon (Salmo salar) underyearlings (Abstract). Aquaculture. 294.
- (1993). Growth and Parr-smolt tranformation of Atlantic Salmon (Salmo salar L.) under different light intensities and subsequent survival and growth in seawater. Aquacultural Engineering. 231-243.
- (1992). The potential for development of salinity tolerance in underyearling Atlantic salmon (Salmo salar). Journal of the World Aquaculture Society. 52-55.
- (1992). The potential for development of salinity tolerance in underyearling Atlantic salmon (Salmo salar L.). World Aquaculture. 52-55.
- (1992). Growth and sexual maturation in Atlantic salmon, Salmo salar L. reared in sea cages at two different light regimes. Aquaculture and Fisheries Management. 275-280.
- (1992). First feeding of Atlantic salmon (Salmo salar L.) under different photoperiods. Journal of Applied Ichthyology. 251-256.
- (1992). Advancement and synchronization of ovulation in Atlantic salmon (Salmo salar L.) following injections of LHRH analogue. Aquaculture. 169-175.
- (1991). Growth, parr-smolt transformation and changes in growth hormone of Atlantic salmon (Salmo salar) reared under different photoperiods. Canadian Journal of Fisheries and Aquatic Sciences. 2100-2108.
- (1991). Growth and sexual maturation in Atlantic salmon (Salmo salar) reared in sea cages at two different light regimes. Aquaculture and Fisheries Management. 275-280.
- (1991). First feeding of Atlantic salmon (Salmo salar) under different photoperiods. Journal of Applied Ichthyology. 251-256.
- (1991). Continous light increases growth of Atlantic salmon (Salmo salar) postsmolts in seacages. Aquaculture. 281-287.
- (1990). First feeding of Atlantic salmon, Salmo salar L., under different photoperiods and light intensities. Aquaculture and Fisheries Management. 435-441.
- (1989). The effect of spectral composition on growth and smoltification in Atlantic salmon (Salmo salar) and subsequent growth i sea cages. Aquaculture. 155-162.
- (1989). Effects of tank colour on growth and smoltification of Atlantic salmon (Salmo salar L.). Aquaculture. 379-386.
- (2019). The Freshwater Institute & CtrlAQUA, Norway: A multi-year research collaboration on RAS-based Atlantic salmon production. .
- (2018). Adaptive immune responses of Atlantic salmon (Salmo salar L.) to stimulation by Salmonid alphavirus subtype 3 infection or vaccination.
- (2017). The effects of photoperiod on Atlantic salmon post-smolt in freshwater closed-containment systems.
- (2017). My skin don't lie. .
- (2017). Korleis få professorar med på ein kollegial SoTL-kultur?
- (2016). Semi-closed sea systems for postsmolt Atlantic salmon. (Abstract ID 42049).
- (2016). SAV3 infection and shedding rate is dose dependent.
- (2016). Osmoregulatory changes in sexually maturing Atlantic salmon post-smolt and adults.
- (2016). Effects of androgens on osmoregulatory mechanisms in Atlantic salmon (Salmo salar L.).
- (2016). Androgens up-regulate freshwater ion transporters in seawater acclimated Atlantic salmon (Salmo salar).
- (2015). Semi-closed sea systems for postsmolt Atlantic salmon. Conference Abstract p. 123.
- (2015). Physiological and immunological parameters affecting salmonid alphavirus infection in Atlantic salmon at post-smolt stages.
- (2015). Immunological parameters correlating with viraemic period of salmonid alphavirus infection in Atlantic Salmon post-smolts.
- (2015). Atlantic salmon, salmo salar I., post-smolts are more susceptible to salmonid alphavirus infection during the initial phase after seawater transfer.
- (2015). Atlantic salmon, Salmon salar1, post-smolts are more susceptible to salmon alphavirus infection during the initial phase after sea water transfer.
- (2015). Atlantic salmon post-smolts in RAS: effects of salinity, exercise and timing of seawater transfer on performance, physiology and welfare. In Book of abstracts, p. 39.
- (2015). Atlantic salmon post-smolts in RAS: Effects of salinity, exercise and timing of seawater transfer on performance, physiology and welfare.
- (2014). Tidlig kjønnsmodning påvirker laksens evne til å regulere vann- og salt balansen i sjø - et velferdsproblem?
- (2014). The effects of increased CO2 on postsmolt Atlantic salmon Salmo salar growth, physiology and stress responsiveness.
- (2014). Saltholdighet og fotoperiode påvirker postsmoltmodning og genuttrykk hos laksehanner.
- (2014). Nutrient transport in the Atlantic salmon (Salmo salar) is affected by environmental salinity.
- (2014). MitSAV: Mitigating the challenges in the Atlantic salmon aquaculture caused by salmonid alphavirus by unveiling the underlying immune mechanisms.
- (2014). Effects of androgens and cortisol on gill ion transporters and osmoregulatory capacity in atlantic salmon.
- (2014). EXPLORING PHYSIOLOGICAL FUNCTIONS OF THE PLEIOTROPHIC MELANOCORTIN ALPHA-MSH IN ATLANTIC SALMON.
- (2013). Effects of salinity and exercise on Atlantic salmon postsmolts reared in land-based RAS.
- (2012). Økende temperaturer fremskynder smoltutvikling hos Vosso laks. Resultat fra prosjektet: "The smolt probe - novel tools for assessment of smolt quality and marine performance in Atlantic salmon (SmoltProbe)".
- (2009). Salmon lice induced mortality of Atlantic salmon postsmolts experiencing a previous history of poor water quality in freshwater.
- (2009). Do exposure to saline water and intensive rearing during smoltification affect performance and disease resistance in Atlantic salmon?
- (2009). Atlantic salmon smolt recovery rates following short term aluminum exposure.
- (2008). Exploring the roles of leptin in fish.
- (2006). Long-term separate and combined effects of environmental hypercapnia and hyperoxia in Atlantic Salmon Salmo salar L.smolts.
- (2006). Long-term effects of reduced water flow, increased carbon dioxide and reduced pH(AI) on Atlantic Salmon welfare Salmo salar L.
- (2005). Status og utfordringer innen vannkvalitet og vannbehandling for ulike livsstadier av laks ved intensivt oppdrett.
- (2005). Salmon lice or suboptimal water quality ? reasons for reduced postsmolt survival?
- (2005). Intensive rearing and welfare in juvenile Atlantic salmon.
- (2005). Exposure to acid water and aluminium reduces Atlantic salmon postsmolt survival.
- (2004). O2-overmetning. Fysiologi, helse og prestasjoner i sjøperioden.
- (2004). Intensivt oppdrett av laksesmolt: konsekvenser for laksens prestasjoner i ferskvannsfasen og etter overføring til sjø.
- (2004). Forsuring eller lakselus: effekter på postsmoltoverlevelse.
- (2002). Smolting in anadromous and landlocked strains of Atlantic salmon (Salmo salar).
- (2002). Effect of different dietary lipids on Atlantic salmon (Salmo salar) growth, fatty acid composition of Gills and Gill Na+, K+-ATPase Activity.
- (2002). EFFECTS OF PHOTOPERIOD, TEMPERATURE AND GnRHa TREATMENT ON THE TIMING OF SPAWNING AND EGG AND SPERM QUALITY IN ATLANTIC SALMON (SALMO SALAR L.).
- (2000). Three forms of GnRH in the brain and pituitary of the turbot, Scophthalmus maximus: immunological characterization and seasonal variation. Comp. Biochem. Physiol. 00, 000-000.
- (1996). Lysregimet påvirker vekst hos ung kveite (Hippoglossus hippoglossus) i oppdrett.
- (1994). Photoperiodic regulation of plasma growth hormone levels during parr-smolt transformation of Atlantic salmon: implications for hypoosmoregulatory ability and growth.
- (1994). Photoperiod and temperature control of salinity tolerance, gill NA+, K+-ATPase activity and seawater performance underyearling Atlantic salmon (salmo salar L.).
- (1993). Sammenheng mellom smoltkvalitet og immunsvar hos laks etter vaksinasjon med Vibrio salmonicida.
- (1993). Growth of O%+ halibut related to temperature, light period and feeding regime.
- (1992). The importance of fish size and photoperiod on growth in Atlantic salmon (Salmo salar) 0+ smolts.
- (1992). Smolting in underyearling Atlantic salmon (Salmo salar).
- (1992). Photoperiodic effects on growth, gonadal development and flesh colouring in Atlantic salmon (Salmo salar).
- (1992). Photoperiodic control of sexual maturation in Atlantic salmon.
- (1992). Parr-smolt transformation of Atlantic salmon (Salmo salar) at different water temperatures.
- (1992). Growth and parr-smolt transformation of individually tagged Atlantic salmon (Salmo salar L.) transferred between continuous light and simulated natural photoperiod.
- (1992). Effects of reduced daylength on growth and bimodality in Atlantic salmon (Salmo salar L.).
- (1992). Effects of photoperiod and temperature on growth and parr-smolt transformation in Atlantic salmon (Salmo salar L.) and subsequent post-smolt performance in seawater.
- (1991). Photoperiodic control of plasma growth hormone levels and sexual maturation of adult Atlantic salmon.
- (1991). Photoperiodic control of growth, incidence of sexual maturation and ovulation in adult Atlantic salmon.
- (1991). Growth and sexual maturation of Atlantic salmon under two temperature regimes in floating enclosures.
- (1991). Growth and sexual maturation in reared Atlantic salmon (Salmo salar).
- (1991). Factors affecting the quality of Atlantic salmon smolts for use in sea ranching.
- (1991). Effects of photoperiod on plasma GtH 1 and GtH 2 levels during sexual maturation in Atlantic salmon.
- (1991). Effect of photoperiod during parr-smolt transformation on sexual maturation in Atlantic salmon.
- (1991). Consequences of seawater acclimation on the survival of Atlantic salmon (Salmo Salar L.) smolts in the presence of cod (Gadus morhua L.).
- (2005). Intensive rearing and welfare in juvenile Atlantic salmon. ?. 449-450.
More information in national current research information system (CRIStin)
Selected publications (2013-2017)
2017
Imsland, A.K.D., Roth, B., Fjelldal, P.G., Stefansson, S.O., Handeland, S. and Mikalsen, B., 2017. The effect of continuous light at low temperatures on growth in Atlantic salmon reared in commercial size sea pens. Aquaculture, 479, 645-651. DOI: 10.1016/j.aquaculture.2017.07.014
Moore L. J., Nilsen T. O., Jarungsriapisit J., Fjelldal P. G., Stefansson S. O., Taranger G. L. and Patel, S., 2017. Triploid Atlantic salmon (Salmo salar L.) post-smolts accumulate prevalence more slowly than diploid salmon following bath challenge with salmonid alphavirus subtype 3. PLoS ONE 12(4): e0175468. https://doi.org/10.1371/journal.pone.0175468
Moore, L. J., Jarungsriapisit, J., Nilsen, T. O., Stefansson, S., Taranger, G. L., Secombes, C. J., Morton, H. C. and Patel, S., 2017. Immune gene profiles in Atlantic salmon (Salmo salar L.) post-smolts infected with SAV3 by bath-challenge show a delayed response and lower levels of gene transcription compared to injected fish. Fish and Shellfish Immunology, 62, 320-331.
Calabrese, S., T.O. Nilsen, J. Kolarevic, L.O.E. Ebbesson, C. Pedrosa, S. Fivelstad, C. Hosfeld, S.O. Stefansson, B.F. Terjesen, H. Takle, C.I.M. Martins, H. Sveier, F. Mathisen, A.K. Imsland and S.O. Handeland, 2017. Stocking density limits for post-smolt Atlantic salmon (Salmo salar L.) with emphasis on production performance and welfare. Aquaculture, 468, 363-370. DOI: 10.1016/j.aquaculture.2016.10.041
2016
Imsland, A. K., K. Pettersen and S. O. Stefansson, 2016. Growth and smoltification of three Norwegian strains of Atlantic salmon reared under different thermal regimes. River Res. Applic., 32, 1800–1807. DOI: 10.1002/rra.3011
Døskeland, I., Imsland, A. K., Fjelldal, P. G., Stefansson, S. O., Roth, B., Eriksen, K. E., Mikalsen, B. and Handeland, S. O., 2016. The effect of low temperatures and photoperiods on growth and vertebra morphometry in Atlantic salmon. Aquacult. Int., 24,1421–1434. DOI 10.1007/s10499-016-9999-7
Jarungsriapisit, J., L. J. Moore, G. L. Taranger, T. O. Nilsen, H. C. Morton, I. U. Fiksdal, S. Stefansson, P. G. Fjelldal, Ø. Evensen and S. Patel, 2016. Atlantic salmon (Salmo salar L.) post-smolts challenged two or nine weeks after seawater-transfer show differences in their susceptibility to salmonid alphavirus subtype 3 (SAV3) Virology Journal, 13 Article Number 66 DOI: 10.1186/s12985-016-0520-8
Vikingstad, E., Andersson, E., Hansen, T. J., Norberg, B., Mayer, I., Stefansson, S. O., Fjelldal, P. G. and Taranger, G. L., 2016. Effects of temperature on the final stages of sexual maturation in Atlantic salmon (Salmo salar L.). Fish Physiol. Biochem. 42,.895-907. DOI: 10.1007/s10695-015-0183-1
Angotzi, A.R, S. O. Stefansson, T. O. Nilsen, J. I. Øvrebø, E. Andersson, G. L. Taranger and I. Rønnestad, 2016. Identification of a novel leptin receptor duplicate in Atlantic salmon: Expression analyses in different life stages and in response to feeding status. Gen. Comp. Endocrinol., 235, 108-119, http://dx.doi.org/10.1016/j.ygcen.2016.06.004
Sveen, L. R., Timmerhaus, G., Torgersen, J. S., Ytteborg, E., Jørgensen, S. M., Handeland, S., Stefansson, S. O., Nilsen, T. O., Calabrese, S., Ebbesson, L., Terjesen, B. F. and Takle, H., 2016, Impact of fish density and specific water flow on skin properties in Atlantic salmon (Salmo salar L.) post-smolts, Aquaculture, 00, 000-000, doi: 10.1016/j.aquaculture.2016.08.012
2015
Salmerón, C., Johansson, M., Asaada,M., Angotzi, A.R., Rønnestad, I., Stefansson, S. O., Jönsson, E., Björnsson, B. Th., Gutiérrez, J., Navarro, I. and Capilla, E., 2015. Roles of leptin and ghrelin in adipogenesis and lipid metabolism of rainbow trout adipocytes in vitro. Comp. Biochem. Physiol., A 188, 40-48. doi:org/10.1016/j.cbpa.2015.06.017
2014
Sundh, H., Nilsen, T.O., Lindström, J., Hasselberg-Frank, L., Stefansson, S.O., McCormick, S.D., Sundell, K. (2014). Development of intestinal transport mechanisms during smoltification in Atlantic salmon smoltification and acclimation to seawater. J. Fish Biol. 85, 1227-1252. doi:10.1111/jfb.12531
Handeland, S.O., Imsland, A.K., Nilsen, T.O., Ebbesson, L.O.E., Hosfeld, C.D., Pedrosa, C., Stefansson, S.O., (2014). Osmoregulation in wild Atlantic salmon smolts transferred to seawater at different temperatures. J. Fish Biol., 85, 1163-1176. doi:10.1111/jfb.12481
Einarsdóttir, I.E., Gong, N., Jönsson-Bergman, E., Sundh, H., Hasselberg Frank, L., Nilsen, T.O., Stefansson, S.O., Sundell, K., Björnsson, B.Th., (2014). Plasma growth hormone binding protein levels in Atlantic salmon (Salmo salar L.) during smoltification and seawater transfer. J. Fish Biol. 85, 1279-1296. doi: 10.1111/jfb.12473
Imsland, AK, Handeland, SO and Stefansson, SO, 2014. Photoperiod and temperature effects on growth and maturation of pre- and post-smolt Atlantic salmon. Aquaculture International, 22, 1331-1345. doi: 10.1007/s10499-014-9750-1
Handeland, SO, Imsland, AK, Ebbesson, LOE, Nilsen, TO, Hosfeld, CD, Teien, HC and Stefansson, SO, 2014. Osmoregulation and growth in offspring of wild Atlantic salmon at different temperatures. Env. Biol. Fishes, 97, 285-296. doi: 10.1007/s10641-013-0151-5
2013
Nilsen, TO, LOE Ebbesson, SO Handeland, F Kroglund, B Finstad, AR Angotzi, SO Stefansson, 2013. Atlantic salmon (Salmo salar L.) smolts require more than two weeks to recover from acidic water and aluminum exposure. Aquatic Toxicol., 142, 33-44. doi.org/10.1016/j.aquatox.2013.07.016
Imsland, A. K., Handeland, S. O., Björnsson, B. Th. and Stefansson, S. O., 2013. Long-term effects of photoperiod and temperature and their interaction on growth, gill Na+,K+-ATPase activity, seawater tolerance and plasma growth hormone levels in Atlantic salmon. J. Fish Biol., 83, 1197-1209
Grassie, C., V.A. Braithwaite, J. Nilsson, T.O. Nilsen, H-C. Teien, S.O. Handeland, S.O. Stefansson, V. Tronci, M. Gorissen, G. Flik & L.O.E. Ebbesson, 2013. Aluminum exposure impacts brain plasticity and behavior in Atlantic salmon (Salmo salar). J. Exp. Biol., 216, 3148-3155, doi:10.1242/jeb.083550
Angotzi, A. R., Stefansson, S., Nilsen, T. O., Rathore, R. M. and Rønnestad, I. 2013. Molecular cloning and genomic characterization of novel Leptin-like genes in salmonids provide new insight into the evolution of the Leptin gene family. Gen. Comp. Endocrinol., 187, 48-59, doi: 10.1016/j.ygcen.2013.03.022
Handeland, S. O., Imsland, A. K., Ebbesson, L. O. E., Nilsen, T. O., Hosfeld, C. D., Teien, H. C. and Stefansson, S. O., 2013. Osmoregulation and growth in offspring of wild Atlantic salmon at different temperatures. Environmental Biology of Fishes, 97, 285-296.
Handeland, S. O., Imsland, A. K., Ebbesson, L. O. E., Nilsen, T. O., Hosfeld, C. D., Baeverfjord, G., Espmark, Å., Rosten, T., Skilbrei, O. T., Hansen, T., Gunnarsson, G. S., Breck, O. and Stefansson, S. O., 2013. Low light intensity can reduce Atlantic salmon smolt quality. Aquaculture, 384-387, 19-24. doi: org/10.1016/j.aquaculture.2012.12.016
Handeland, S. O., Imsland, A. K, Björnsson, B. Th., Stefansson, S. O. and Porter, M. 2013. Physiology during smoltification in Atlantic salmon: effect of melatonin implants. Fish Physiol. Biochem., 39, 1079-1088. doi: 10.1007/s10695-012-9765-3
Urke, H.A., Kristensen, T., Arnekleiv, J.V., Haugen, T.O., Kjærstad, G., Stefansson, S.O., Ebbesson, L.O.E. and Nilsen, T.O., 2013. Seawater tolerance and post-smolt migration of wild Atlantic salmon Salmo salar × brown trout S. trutta hybrid smolts, J. Fish Biol., 82, 206–227, doi:10.1111/j.1095-8649.2012.03481.x
National projects (Research Council of Norway)
NFFR V.701.146 'Miljøforholdene i settefiskproduksjon', 1986 - 1988
NTNF HB 23.22395 'Lys i fiskeoppdrett', 1987 - 1989
NFFR V.701.211 'Halvtårssmolt', 1988 - 1990
NFFR V.701.236 'Lys i matfiskproduksjon', 1989 - 1991
NFFR 1401-701.321 'Smoltkvalitet og helse', 1991 - 1993
NFR 108848 ‘Smoltkvalitet, sesonguavhengig utsetting og tidlig sjøvannsfase’, 1995 - 1998
NFR 108098 ‘Postsmoltøkologi – trofisk nivå og vandringer’, 1995 - 1997
NFR 110993 ‘Miljømessig og arvelig regulering av vekst hos ung kveite i oppdrett’, 1996 - 1999
NFR 122845 ‘Det arvelige grunnlaget for smoltifisering hos oppdrettslaks og villaks – en grunnleggende studie’, 1998 - 2000
NFR 133964 ’Vannbehov og vannkvalitet under intensiv smoltproduksjon. Hva er grenseverdiene og hvordan unngå skadelige effekter av lav vanngjennomstrømming, lav pH og høg CO2 under intensiv smoltproduksjon?’, 2000 – 2002
NFR 133937 ‘Control of age at first sexual maturity (puberty) in farmed Atlantic salmon’, 2000 – 2002
NFR 149790 'Distribution and ecology of post-smolt and salmon at sea', 2002 - 2005
NFR 149796 ‘The importance of early marine feeding on the growth and survival of Atlantic salmon post-smolt in Norwegian fjords’, 2002 - 2006
NFR 153230 ‘Environmental control of smolt brain development’, 2003 – 2005
NFR 165331 ‘Environmental impact and molecular regulation of the osmoregulatory gill during parr-smolt transformation in Atlantic salmon’, 2005 – 2007
NFR 163429 ‘Optimisation of harvested halibut by lowering maturation, increasing growth and improving flesh quality’ 2005 – 2007
NFR 165239 ‘Sjøvannsinnblanding i intensiv smoltproduksjon: effekter på helse, velferd og risiko for vintersår’, 2005 – 2007
NFR 172504 ’Nitric oxide regulation of development in fish: emphasis on the light-brain-pituitary axis and gill during salmon smoltification’, 2006 - 2008
NFR 1722630 ‘Progressive, modern production of juvenile Atlantic cod’ (PROCOD), 2006 - 2009
NFR 172514 ‘Moderne settefiskproduksjon av laks - normal utvikling og velferd i intensive oppdrettsystemer’, 2006 - 2009
NFR 184997 ‘Optimal smolt production and post smolt performance in the High North - Seawater intermixing, low temperatures and intensive rearing’ (NORSMOLT), 2008 – 2011
NFR 190469 ‘Fish welfare assessment through brain function: Impacts of environmental stress on brain plasticity and behaviour’, 2009 – 2011
NFR 199800 ‘The smolt probe - novel tools for assessment of smolt quality and marine performance in Atlantic salmon’, 2010 – 2012
NFR 199518 ‘Early puberty in salmon males in seawater - causes and consequences for hypoosmoregulatory ability and welfare’, 2010 – 2013
NFR 199482 ‘Cod Development CODE’, 2010 – 2013
NFR 222215 ’The smolt brain model: Unraveling nature´s regulation of neural plasticity’, 2013 – 2015
NFR 233870 ‘Welfare of Atlantic salmon postsmolts in closed-containment production systems, using a function-based approach’ (SalmoFutura), 2014-2016
NFR 256199 ‘Optimal bruk av rognkjeks til avlusing av oppdrettslaks, LUSINFE’R, 2016 – 2020
NFR 269043 ‘Kontroll av kjønnsmodning hos rognkjeks, STAMINA’, 2017 – 2021
National strategic programs (Research Council of Norway)
114646 ‘Teori- og eksperimentbasert studie av rekruttering hos marin fisk’, 1995 - 1998
136260 ‘Smolting in Atlantic salmon: the interrelationship between physiological, endocrine and immunological changes, and their genetic regulation’, 2000 - 2006
Other national programs (Fiskeri og havbruksnæringens forskningsfond, FHF)
FHF/NFR 155157 'Vannkvalitet – smoltkvalitet', 2003 – 2005
International projects
Nordisk Industrifond 88.107 'Helårlig produksjon av atlantisk laks ved styrt kjønnsmodning og smoltifisering', 1988 - 1991
EU, FAIR-CT96-1410 ‘Environmental and neuroendocrine control mechanisms in finfish reproduction and their application in broodstock management’, 1997 - 1999
EU, FAIR-CT97-3544 ‘A multidisciplinary evaluation and optimization of the production characteristics of different strains of commercially cultured flatfish’, 1998 - 2000
Wallenberg foundation, 1997 - 2001
‘Aquaculture and Fisheries Development: Basic Research on Finfish Reproduction and Growth Enabling the Intensification of Aquaculture’
‘The Multiple Gonadotropin-Releasing Hormones (GnRH) of Fish: Functions and Potential Use for Spawning Manipulations in Commercially and Ecologically Important Fish Species’
EU, QoL, QLRT-2001-01801 ‘Photoperiod control of puberty in farmed fish: Development of new techniques and research into underlying physiological mechanisms’, 2002 - 2005
EU, LSF ‘Marine Pelagic Food Chain Research’, (Contract No ERBFMGECT950013)
Research collaborators/topics:
Professor Niall Bromage, University of Stirling, Skottland
Temperature effects on melatonin in post-smolt Atlantic salmon (Salmo salar), 1997
Professor Sjoerd E. Wendelaar Bonga, Katholieke Universiteit Nijmegen, Nederland
The effect of temperature on growth performance and stress resistance in two generations of turbot (Scophthalmus maximus Rafinesque), 1997
Influence of temperature and feeding ration on the bioenergetics of juvenile turbot (Scophthalmus maximus) and Atlantic halibut (Hippoglossus hippoglossus), 1998
Dr. Richard FitzGerald, University College Cork, Irland
The effect of different initial size distribution on the growth of Atlantic halibut (Hippoglossus hippoglossus L.), 1998
Dr. Ewa Kulczykowska, Marine Biology Center of Polish Academy of Sciences, Gdynia, Polen
Melatonin – Arginine vasotocin relationship during development of Salmo salar, 2001
EU, Improving Human Potential, Access to Research Infrastructure, Institute of Aquaculture, University of Stirling, Scotland, Contract HPRI-CT-1999-00106
Professor Niall Bromage
Endocrine regulation of growth and food intake in salmon smolts, 2000/2001
EU, FP7-KBBE-2007-2A, 222719, LIFECYCLE—Building a biological knowledge-base on fish lifecycles for competitive, sustainable European aquaculture (01.03.2009 – 28.02.2013)
CentreCentre for research based innovation (SFI): Centre for Research-based Innovations in Controlled-environment Aquaculture (CtrlAQUA), 2015 – 2023. http://ctrlaqua.no/
Cand. scient., Dept. of Fisheries Biology, University of Bergen (UiB), 25.06.1986
Thesis: Lysperiodens virkning på vekst og smoltifisering hos laks (Salmo salar L.)
Dr. scient., Dept. of Fisheries and Marine Biology, UiB, 10.12.1990
Thesis: The influence of light on growth and parr-smolt transformation of Atlantic salmon (Salmo salar L.): Effects of spectral composition, light intensity and photoperiod