- E-mailhans.skaug@uib.no
- Phone+47 55 58 48 61+47 469 45 448
- Visitor AddressRealfagbygget, Allegaten 41, 4 etg. sør.BergenRoom4A17d
- Postal AddressPostboks 78035020 Bergen
Biostatistics
I am interested in applications of statistics and probability to biology, especially to marine ecology. I have developed methods for line transect surveys and CKMR (close-kin mark recapture) methods in particular. Key reference: Bravington, MV, Skaug, HJ, Anderson, EC (2016). Close-kin Mark-Recapture. Statistical Science.
Computational statistics
My opinion for a long time has been that Statisticians should have Automatic Differentiation (AD) in their toolbox. My conjecture has been that a combination of AD and the Laplace approximation is a powerful platform for fitting models with continuous latent variables. Software projects in which this idea is implemented are:
- ADMB: http://admb-project.org/
Artificial intelligence (AI)
Witnessing the recent breakthroughs in the field, my research interests have turned towards AI. Techniques such as variational autoencoders and diffusion models are probabilistic in nature, and Backpropagation in Deep Learning is the same computational technique as AD, which I have been working with for 20 years.
Editorial services
co-Editor in chief of Scandinavian Journal of Statistics (2018–2021)
Teaching at UiB (last 5 years)
Short courses / workshops (in the past)
- CKMR (Close-kin mark recapture) course (co-thought) Dalhousie University, Halifax, July 2019.
- TMB (Template Model Builder) course (co-thought) at IMR Bergen, June 2019.
- Modelling in TMB (Template Model Builder). Course (co-thought) at IMR Bergen, June 2018.
- An introduction to fish stock assessment in TMB (Template Model Builder). Course given Oct. 2017 in La Jolla
From CRIStin (Current research information system in Norway):
- (2023). A TMB Approach to Study Spatial Variation in Weather-Generated Claims in Insurance. SN Operations Research Forum.
- (2022). The heritability of BMI varies across the range of BMI—a heritability curve analysis in a twin cohort. International Journal of Obesity. 1786-1791.
- (2022). Mixed Frequency Data in a (S,s) Pricing Model.
- (2022). Evaluating the suitability of close-kin mark-recapture as a demographic modelling tool for a critically endangered elasmobranch population. Evolutionary Applications. 1-13.
- (2021). Variation in use of Caesarean section in Norway: An application of spatio-temporal Gaussian random fields. Scandinavian Journal of Public Health. 8 pages.
- (2021). The Sibling Distribution for Multivariate Life Time Data. Sankhya B, The Indian Journal of Statistics. 340-363.
- (2021). Report of the NAMMCO-ICES Workshop on Seal Modelling (WKSEALS 2020). NAMMCO scientific publications.
- (2021). Measurement error model for the Norwegian common minke whale (Balaenoptera acutorostrata acutorostrata) surveys 2008-2013. Journal of Cetacean Research and Management. 16 pages.
- (2021). Epistemic uncertainty quantification in deep learning classification by the Delta method. Neural Networks. 164-176.
- (2021). Considering sampling bias in close-kin mark–recapture abundance estimates of Atlantic salmon. Ecology and Evolution.
- (2021). Consideration of measurement errors for the Norwegian common minke whale (Balaenoptera acutorostrata acutorostrata) surveys. Journal of Cetacean Research and Management. 1-16.
- (2020). Heritability curves: A local measure of heritability in family models. Statistics in Medicine. 1357-1382.
- (2020). Experimental design for parameter estimation in steady-state linear models of metabolic networks. Mathematical Biosciences.
- (2020). Estimation and identifiability of kinetic parameters in dynamical models of biochemical reaction networks.
- (2020). A comparison of Monte Carlo sampling methods for metabolic network models. PLOS ONE. 1-24.
- (2019). dCOD 1.0: DECODING THE SYSTEMS TOXICOLOGY OF ATLANTIC COD (GADUS MORHUA) – HIGHLIGHTS SO FAR.
- (2019). dCOD 1.0: DECODING THE SYSTEMS TOXICOLOGY OF ATLANTIC COD (GADUS MORHUA) – HIGHLIGHTS SO FAR.
- (2019). Validation of close‐kin mark–recapture (CKMR) methods for estimating population abundance. Methods in Ecology and Evolution. 1445-1453.
- (2019). Stochastic data assimilation of observations with a detection limit.
- (2019). Integration by differentiation; almost a free lunch.
- (2019). Efficient Computation of Hessian Matrices in TensorFlow . arXiv.org.
- (2019). Editorial. Scandinavian Journal of Statistics. 1-1.
- (2019). Drivers of the summer-distribution of Northeast Atlantic mackerel (Scomber scombrus) in the Nordic Seas from 2011 to 2017; a Bayesian hierarchical modelling approach. ICES Journal of Marine Science. 530-548.
- (2019). Contaminant accumulation and biological responses in Atlantic cod (Gadus morhua) caged at a capped waste disposal site in Kollevåg, Western Norway. Marine Environmental Research. 39-51.
- (2019). Close-Kin Mark-Recapture (CKMR).
- (2019). Close-Kin Mark-Recapture (CKMR).
- (2019). Close kin mark recapture in fisheries .
- (2019). Bayesian flux balance analysis of metabolic pathways.
- (2019). Bayesian and frequentist computation with application to data from the Medical Birth Registry of Norway.
- (2018). dCOD 1.0: DECODING THE SYSTEMS TOXICOLOGY OF ATLANTIC COD (GADUS MORHUA).
- (2018). Model Reduction for Tracer Transport and Applications.
- (2018). Mathematical methods for detection and localization of CO2 leaks.
- (2018). Machine Learning in CO2 leak detection.
- (2018). Experimental design for estimation of first order kinetic parameters in steady-state metabolic networks.
- (2018). Ecotoxicological responses in Atlantic cod (Gadus morhua) after caging at a capped waste disposal site in Kollevåg, Western Norway .
- (2018). Ecotoxicological responses in Atlantic cod (Gadus morhua) after caging at a capped waste disposal site in Kollevåg, Western Norway .
- (2018). Combining Models and Machine Learning Techniques to Design Leak Detection Monitoring .
- (2018). Bayesian flux balance analysis; concepts and computation.
- (2017). glmmTMB balances speed and flexibility among packages for zero-inflated generalized linear mixed modeling. The R Journal. 378-400.
- (2017). dCod 1.0: decoding the systems toxicology of Atlantic cod (Gadus morhua).
- (2017). dCod 1.0 decoding the systems toxicology of Atlantic cod.
- (2017). Whole genome resequencing reveals diagnostic markers for investigating global migration and hybridization between minke whale species. BMC Genomics. 1-11.
- (2017). The parent-offspring probability when sampling age-structured populations. Theoretical Population Biology. 20-26.
- (2017). TMB: a flexible framework for developing mixed model software in R.
- (2017). Structural identifiability of kinetic parameters.
- (2017). Structural identifiability of kinetic parameters.
- (2017). PRECISION-CUT LIVER SLICES (PCLS) FOR TOXICOGENOMIC STUDIES IN ATLANTIC COD (Gadus morhua): THE dCOD 1.0 PROJECT.
- (2017). On the application of improved symplectic integrators in Hamiltonian Monte Carlo. Communications in Statistics - Simulation and Computation. 500-509.
- (2017). Mathematical modeling of regulatory interactions in the fatty acid synthesis pathway.
- (2017). Mathematical modeling of regulatory interactions in the fatty acid synthesis pathway.
- (2017). Mathematical modeling of regulatory interactions in the fatty acid synthesis pathway.
- (2017). Bayes’ theorem as the fundament to design monitoring programs.
- (2017). Assessment of machine learning methods as a tool in detecting leakages.
- (2017). Assessment of Topological Data Analysis and Machine Learning Technologies as Tools for Seep Detection.
- (2016). Trade-offs between accuracy and interpretability in von Bertalanffy random-effects models of growth. Ecological Applications. 1535-1552.
- (2016). TMB: Automatic differentiation and laplace approximation. Journal of Statistical Software. 1-21.
- (2016). Likelihood Estimation of Jump-Diffusions: Extensions from Diffusions to Jump-Diffusions, Implementation with Automatic Differentiation, and Applications.
- (2016). Close-kin mark-recapture. Statistical Science. 259-274.
- (2016). Bayes’ theorem as the fundament to design monitoring programs.
- (2015). The importance of spatial models for estimating the strength of density dependence. Ecology. 1202-1212.
- (2015). Spatial factor analysis: A new tool for estimating joint species distributions and correlations in species range. Methods in Ecology and Evolution. 627-637.
- (2015). Panel data model with mixed frequency – an application to producers' pricing behaviour.
- (2015). Measurement error model for the Norwegian minke whale surveys 2008-2013.
- (2015). Geostatistical delta-generalized linear mixed models improve precision for estimated abundance indices for West Coast groundfishes. ICES Journal of Marine Science. 1297-1310.
- (2015). Fitting state-space models to seal populations with scarce data. ICES Journal of Marine Science. 1462-1469.
- (2015). Distance sampling with a random scale detection function. Environmental and Ecological Statistics. 725-737.
- (2015). Data assimilation by use of the iterative ensemble smoother for 2D facies models. SPE Journal. 169-185.
- (2015). DNA-register for vågehval: en skattekiste for studier av hvalbiologi. Fisken og Havet, Særnummer. 108-109.
- (2015). Bandwidth selection in pre-smoothed particle filters. Statistics and computing. 16 pages.
- (2015). Abundance estimates of common minke whales in the Northeast Atlantic base don survey data collected over the period 2008-2013.
- (2014). Unødvendig med fem fangstområder for vågehval. Forskning.no.
- (2014). Preliminary abundance estimates of common minke whales in the Northeast Atlantic based on survey data collected over the period 2008-2013.
- (2014). Next-generation sequencing for molecular ecology: a caveat regarding pooled samples. Molecular Ecology. 502-512.
- (2014). Maximum likelihood estimation of partially observed diffusion models. Journal of Econometrics. 73-80.
- (2014). Investigating population genetic structure in a highly mobile marine organism: The minke whale balaenoptera acutorostrata acutorostrata in the north east atlantic. PLOS ONE.
- (2014). Estimation of production rates with transient well-flow modeling and the auxiliary particle filter. SPE Journal. 172-180.
- (2014). Determining individual variation in growth and its implication for life-history and population processes using the Empirical Bayes Method. PLoS Computational Biology.
- (2014). Bandwidth Selection In Pre-Smoothed Particle Filters.
- (2014). A flexible and automated likelihood based framework for inference in stochastic volatility models. Computational Statistics & Data Analysis. 642-654.
- (2013). Strategies for fitting nonlinear ecological models in R, AD Model Builder, and BUGS. Methods in Ecology and Evolution. 501-512.
- (2013). Markov-modulated nonhomogeneous poisson processes for modeling detections in surveys of marine mammal abundance. Journal of the American Statistical Association. 840-851.
- (2013). Hybrids between common and Antarctic minke whales are fertile and can back-cross. BMC Genetics. 11 pages.
- (2013). Estimating genotyping error rates from parent-offspring dyads. Statistics and Probability Letters. 812-819.
- (2012). The Norwegian minke whale DNA register: a data base monitoring commercial harvest and trade of whale products. Fish and Fisheries. 313-332.
- (2012). Numerical comparison of Ensemble Kalman Filter and Randomized maximum Likelihood.
- (2012). Good ethics or political and cultural censoring in science? ICES Journal of Marine Science. 493-497.
- (2012). Fitting general stochastic volatility models using Laplace accelerated sequential importance sampling. Computational Statistics & Data Analysis. 3105-3119.
- (2012). Filtering with state space localized Kalman gain. Physica D : Non-linear phenomena. 1123-1135.
- (2012). Data Assimilation Using the Iterative Ensemble Smoother for 2-D Categorical Models.
- (2012). Comparing the adaptive Gaussian mixture filter with the ensemble Kalman filter on synthetic reservoir models. Computational Geosciences. 467-482.
- (2012). Abundance estimation of long-diving animals using line transect methods. Biometrics. 504-513.
- (2012). AD Model Builder: using automatic differentiation for statistical inference of highly parameterized complex nonlinear models. Optimization Methods and Software. 233-249.
- (2011). Quantifying Monte Carlo Uncertainty in the Ensemble Kalman Filter. SPE Journal. 172-182.
- (2011). Genotyping errors in a calibrated DNA register: implications for identification of individuals. BMC Genetics. 10 pages.
- (2011). Bridging the ensemble Kalman filter and particle filters: the adaptive Gaussian mixture filter. Computational Geosciences. 293-305.
- (2010). Simulated maximum likelihood estimation of continuous time stochastic volatility models. 25 pages.
- (2010). Migration of Antarctic Minke Whales to the Arctic. PLOS ONE.
- (2010). Maximum Simulated Likelihood Methods and Applications. Emerald Group Publishing Limited.
- (2010). Hva kan et register over vågehvalens DNA fortelle om adferd og biologi? Ottar. 10-15.
- (2010). Global occurrence trajectories of microfossils: environmental volatility and the rise and fall of individual species. Paleobiology. 224-252.
- (2010). Estimating genetic architectures from artificial-selection responses: A random-effect framework. Theoretical Population Biology. 119-130.
- (2010). Detecting dyads of related individuals in large collections of DNA-profiles by controlling the false discovery rate. Molecular Ecology Resources. 693-700.
- (2010). Asymptotic bias of the hazard probability model under model mis-specification. Journal of Cetacean Research and Management. 249-252.
- (2010). A method for defining management units based on genetically determined close relatives. ICES Journal of Marine Science. 551-558.
- (2008). Genetic analyses reveal promiscuous mating in female minke whales, Balaenoptera acutorostrata. Journal of Cetacean Research and Management. 249-251.
- (2008). Fitting mixed-effects models when data are left truncated. Insurance, Mathematics & Economics. 121-133.
- (2008). Estimating distributions of correlated individual life-history traits from mark-recapture data.
- (2008). Building and Fitting Non-Gaussian Latent Variable Models via the Moment-Generating Function. Scandinavian Journal of Statistics. 664-676.
- (2007). Historical population assessment of Barents Sea harp sea's (Pagophilus groenlandicus). ICES Journal of Marine Science. 1356-1365.
- (2006). On Curvature and Separability in Unconstrained Optimisation.
- (2006). Markov modulated Poisson processes for clustered line transect data. Environmental and Ecological Statistics. 199-211.
- (2006). DNA registers of legally obtained wildlife and derived products as means to identify illegal takes. Conservation Biology. 1284-1293.
- (2006). Automatic Approximation of the Marginal Likelihood in non-Gaussian Hierarchical Models. Computational Statistics & Data Analysis. 699-709.
- (2006). An integrated-likelihood method for estimating genetic differentiation between populations. Genetics. 2073-2082.
- (2005). Genetic tagging of males in North Atlantic minke whales through comparison of mother and fetus DNA-profiles. Journal of Cetacean Research and Management. 113-117.
- (2005). Genetic tagging of male North Atlantic minke whales through comparison of maternal and foetal DNA-profiles. Journal of Cetacean Research and Management. 113-117.
- (2005). Disentangling the effects of capture efficiency and population abundance on catch data using random effects models. ICES Journal of Marine Science. 1543-1555.
- (2004). Abundance of minke whales (Balaenoptera acutorostrata) in the Northeastern Atlantic. Canadian Journal of Fisheries and Aquatic Sciences. 870-886.
- (2002). Automatic differentiation to facilitate maximum likelihood estimation in nonlinear random effects models. Journal of Computational And Graphical Statistics (JCGS). 458-470.
- (2001). Allele-sharing methods for estimation of population size. Biometrics. 750-756.
- (2000). Towards a statistical model for the uncertainty in fisheries data for stock asssessment in the Barents Sea. SAMBA/06/00. SAMBA/06/00. .
- (1999). Simulated likelihood methods for complex double-platform line transect surveys. Biometrics. 678-687.
- (1999). Hazard models for line transect surveys with independent observers. Biometrics. 29-36.
- (1997). Do Minke Whales (Balaenoptera acutorostrara) Exhibit Particular Prey Preferences? Journal of Northwest Atlantic Fishery Science. 91-104.
- (1997). Abundance of northeastern Atlantic minke whales, estimates for 1989 and 1995. Rep. int. Whal. Commn. 453-483.
- (1996). Perpendicular distance line transect methods based on integrated spatial hazard probability models. Ukjent.
- (1996). NAILS-96, System Design and Program Documentation. Ukjent.
- (1996). NAILS-96 Specification document.
- (1996). Diet and food avaliability for Northeastern Atlantic minke whales, Balaenoptera Acutorostrata. Reports of the International Whaling Commission. 371-382.
- (1996). Abundance of northeastern Atlantic minke whales, estimates for 1989 and 1995. Ukjent.
- (1995). Methods for simulation of stochastic vector-fields in NSPACE. STAT/18/95. STAT/18/95. .
- (1994). Nonparametric tests of serial indepence.
- (1994). Nonparametric functional estimation with application to hypothesis testing.
- (1994). Ikkeparametriske tester for seriell uavhengighet.
- (1993). The limit distribution of the Hoeffding statistic for test of serial independence. Report No.22 Jan. 1993. .
- (1993). A nonparametric test of serial independence based on the empirical distribution function. Biometrika. 591-602.
- (1991). Edgeworth utvikling og bootstrapping i tidsrekke analyse.
More information in national current research information system (CRIStin)