Mollusca Research

Publications (2008 - present)

Main content


Oskars, T. R., and Malaquias, M. A. E. 2022. Systematic revision of the Indo-West Pacific bubble-snails of the genus Haloa (Pilsbry, 1921) (Cephalaspidea: Haminoeidae). Invertebrate Systematicsin press.

Lobo-da-Cunha, A., Bartolomaeus, T. and Malaquias, M. A. E. 2022. Euopisthobranchs. Microscopic Anatomy of Animals, in press.

Hellem, E., and Malaquias, M. A. E. 2022. The taxonomic status of the headshield slug genus Nakamigawaia Kuroda and Habe, 1961 (Gastropoda: Cephalaspidea: Aglajidae), with the description of a new species from the Western Pacific. Journal of Natural Historyin press.

Siegwald, J., Kano, Y., and Malaquias, M. A. E. 2022. A global phylogeny of the deep-sea gastropod family Scaphandridae (Heterobranchia: Cepahalaspidea): redefinition and generic classification. Molecular Phylogenetics and Evolutionin press.

Siegwald, J. and Malaquias, M. A. E. 2022. Revisiting the deep-sea Atlantic gastropod species Scaphander gracilis Watson, 1883: First data on its morphology, systematics, and ecology. Journal of Natural Historyin press.



Ramirez-Llodra, E., Pedersen, T., Dexter, K.F., Hauquier, F., Guilini, K., Mikkelsen, N., Borgersen, G., Van Gyseghem, M., Vanreusel, A. and Vilas, D., 2021. Community structure of deep fjord and shelf benthic fauna receiving different detrital kelp inputs in northern Norway. Deep Sea Research Part I: Oceanographic Research Papers, 168, p.103433. DOI: 10.1016/j.dsr.2020.103433.

Neuhaus, J., Rauch, C., Bakken, T., Picton, B., Pola, M. and Malaquias, M. A. E. 2021. The genus Jorunna (Nudibranchia: Discodorididae) in Europe: a new species and a possible case of incipient speciation. Journal of Molluscan Studies, 87(4), eyab028. DOI: 10.1093/mollus/eyab028

Feliciano, K., Malaquias, M. A. E., Stout, C., Brenzinger, B., Gosliner, T. M., and Valdés, Á. 2021. Molecular and morphological analyses reveal pseudocryptic diversity in Micromelo undatus  (Bruguière, 1792) (Gastropoda: Heterobranchia: Aplustridae). Systematics and Biodiversity, XX: 1-25. https://dx.doi.org/10.1080/14772000.2021.1939458

Araújo, A. K., Pola, M., Malaquias, M. A. E., Ballesteros, M., and Cervera, J. L. 2021. Molecular phylogeny of European Runcinida (Gastropoda, Heterobranchia): the discovery of an unexpected pool of species complex, with special reference to the case of Runcina coronata (Quatrefages, 1844). Zoological Journal of the Linnean Society, XX: 1-28. https://doi.org/10.1093/zoolinnean/zlab041

Martín-Hervás, M. R., Carmona, L., Malaquias, M. A. E., Krug, P. J., Gosliner, T. M. and Cervera, J. L. 2021. A molecular phylogeny of Thuridilla Bergh, 1872 sea slugs (Gastropoda, Sacoglossa) reveals a case of flamboyant and cryptic radiation in the marine realm. Cladistics, 37: 647–676.

Varney, R. M., Brenzinger, B., Malaquias, M. A. E., Meyer, C. P., Schrödl, M. and Kocot, K. 2021.  Assessment of mitochondrial genomes for heterobranch gastropod phylogenetics. BMC Ecology and Evolution, 21(1): 1-14. https://doi.org/10.1186/s12862-020-01728-y



Bharate, M., Apte, D., and Malaquias, M. A. E. 2020. A massive aggregation of the mangrove snail Bakawan rotundata (A. Adams, 1850) (Cephalaspidea: Haminoeidae) in India. Regional Studies in Marine Science, 39: 101406. https://doi.org/10.1016/j.rsma.2020.101406

Sørensen, C. G., Rauch, C., Pola, M. and Malaquias, M. A. E. 2020. Integrative taxonomy reveals a cryptic species of the nudibranch genus Polycera (Polyceridae) in European waters. Journal of the Marine Biological Association of the United Kingdom, 100(5): 733-752. https://doi.org/10.1017/S0025315420000612.

Oskars, T. R. & Malaquias, M. A. E. 2020. The temperate Australasian genus Papawera Oskars and Malaquias, 2019 (Gastropoda: Cephalaspidea: Haminoeidae), with a redescription of P. zelandiae and P. maugeansisJournal of Natural History, 54(21-22): 1343-1362. https://doi.org/10.1080/00222933.2020.1791996.

Oskars, T. R. & Malaquias, M. A. E. 2020. Systematic revision of the Indo-West Pacific colourful bubble-snails of the genus Lamprohaminoea Habe, 1952 (Cephalaspidea : Haminoeidae). Invertebrate Systematics, 34: 727–756. https://doi.org/10.1071/IS20026.

Oskars, T. R. & Malaquias, M. A. E. 2020. Systematic revision of the Indo-West Pacific mangrove-associated snails of the genus Bakawan (Cephalaspidea: Haminoeidae). Journal of Molluscan Studies, 86: 323–341. doi:10.1093/mollus/eyaa012

Lobo-da-Cunha, A., Alves, Â., Oliveira, E., Wranik, W., and Malaquias, M. A. E. 2020. The copulatory apparatus of the marine gastropod Haminella solitaria (Say, 1822) (Heterobranchia: Cephalaspidea) and its phylogenetic relevance. Invertebrate Biology, 139(1): e12281. https://doi.org/10.1111/ivb.12281

Siegwald, J., Pastorino, G., Oskars, T., and Malaquias, M. A. E. 2020. A new species of the deep-sea genus Scaphander (Gastropoda, Cephalaspidea) from the Mar del Plata submarine canyon off Argentina. Bulletin of Marine Sciences, 96(1):111–126. https://doi.org/10.5343/bms.2019.0069



Kocot, K. M., Todt, C., Mikkelsen, N. T., & Halanych, K. M. 2019. Phylogenomics of Aplacophora (Mollusca, Aculifera) and a solenogaster without a foot. Proceedings of the Royal Society B, 286(1902), 20190115. DOI: 10.1098/rspb.2019.0115.

Oskars, T. R. and Malaquias, M. A. E. 2019. A molecular phylogeny of the Indo-West Pacific species of Haloa sensu lato gastropods (Cephalaspidea: Haminoeidae): Tethyan vicariance, generic diversity, and ecological specialization. Molecular Phylogenetics and Evolution, 139: https://doi.org/10.1016/j.ympev.2019.106557

Araujo, A. K., Pola, M., Malaquias, M. A. E., Cervera, J. L. 2019. To be or not to be? What molecules say about Runcina brenkoae Thompson, 1980 1 (Gastropoda: Heterobranchia: Runcinida). Sciencia Marina, 83(3): 223-235. https://doi.org/10.3989/scimar.04907.07A.

Oskars, T. R., Too, C. C., Rees, D., Mikkelsen, P., Willassen, E., Malaquias, M. A. E. 2019. A molecular phylogeny of the gastropod family Haminoeidae sensu lato (Heterobranchia, Cephalaspidea), - a generic revision. Invertebrate Systematics, 33(2): 426-472. https://doi.org/10.1071/IS18051.

Aslam, S., Oskars, T. R., Siddiqui, G., Malaquias, M. A. E. 2019. Beyond shells: First detailed morphological description of the mangrove-associated gastropod Haminoea aff.fusca (A. Adams, 1850) (Cephalaspidea: Haminoeidae), with a COI phylogenetic analysis. Zoosystema, 41(16): 313-326. https://doi.org/10.5252/zoosystema2019v41a16.

Mikkelsen, N. T., Todt, C., Kocot, K. M., Halanych, K. M., Willassen, E. 2019. Molecular phylogeny of Caudofoveata (Mollusca) challenges traditional views.Molecular Phylogenetics and Evolution 132: 138–150. https://doi.org/10.1016/j.ympev.2018.10.037

Moles, J., Avila, C., Malaquias, M. A. E. 2019. Unmasking Antarctic mollusc lineages: novel evidence from philinoid snails (Gastropoda: Cephalaspidea). Cladistics, 35:1–27. https://doi.org/10.1111/cla.12364



Mikkelsen, N. T., Kocot, K. M., Halanych, K. M. 2018. Mitogenomics reveals phylogenetic relationships of caudofoveate aplacophoran molluscs. Molecular Phylogenetics and Evolution127. 429–436. https://doi.org/10.1016/j.ympev.2018.04.031

Mikkelsen, N. T., Todt, C. 2018. One or many? Molecular versus morphological diversity in the aplacophoran Chaetoderma nitidulumLovén, 1844 (Mollusca: Caudofoveata). Journal of Molluscan Studies84. 113–131. https://doi.org/10.1093/mollus/eyy009

Bharate, M., Oskars, T. R., Narayana, S., Ravinesh, R., Kumar, A. B., Malaquias, M. A. E. 2018. Description of a new species of Haminoea (Gastropoda: Cephalaspidea) from India, with an account of the diversity of the genus in the Indo-West Pacific. Journal of Natural History, 52 (37–38): 2437–2456. https://doi.org/10.1080/00222933.2018.1533598

Austin, J., Gosliner, T., Malaquias, M. A. E. 2018. Systematic revision, diversity patterns, and trophic ecology of the tropical Indo-West Pacific sea slug genus PhanerophthalmusA. Adams, 1850 (Cephalaspidea, Haminoeidae). Invertebrate Systematics, 32: 1336–1387. https://doi.org/10.1071/IS17086

Valdés, Á., Breslau, E., Padula, V., Schrödl, M., Camacho, Y., Malaquias, M. A. E., Alexander, J., Bottomley, M., Vital, X., and Gosliner, T. M. 2018. Molecular and morphological systematics of DolabriferaGray, 1847 (Mollusca, Gastropoda, Heterobranchia, Aplysiomorpha). Zoological Journal of the Linnean Society, 184: 31–65. https://doi.org/10.1093/zoolinnean/zlx099

Moles, J., Ávila, C. and Malaquias, M. A. E. 2018. Systematic revision of the Antarctic gastropod family Newnesiidae (Heterobranchia: Cephalaspidea) with the description of a new genus and a new abyssal species. Zoological Journal of the Linnean Society, 183: 763–775. https://doi.org/10.1093/zoolinnean/zlx089

Bharate, M., Oskars, T, Malaquias, M. A. E. 2018. Diniatys callosa (Preston, 1908), new combination name for Haminoea callosafrom the Andaman Islands (India). Journal of Conchology, 43(1): 1–3.

Lobo-da-Cunha, A., Alves, Â., Oliveira, E. and Malaquias, M. 2018. Histological, histochemical and ultrastructural investigation of the male copulatory apparatus of Haminoea navicula(Gastropoda, Cephalaspidea). Journal of Morphology, 279(4): 554–565. https://doi.org/10.1002/jmor.20790 

Zamora-Silva, A. & Malaquias, M. A. E. 2018. Molecular phylogeny of the Aglajidae head-shield sea slugs (Heterobranchia: Cephalaspidea): new evolutionary lineages revealed and proposal of a new classification. Zoological Journal of the Linnean Society, 183(1): 1–51. https://doi.org/10.1093/zoolinnean/zlx064



Oskars,T. R., Mifsud, C., & Malaquias, M. A. E. 2017. Redescription of the Cephalaspidea gastropod Atys jeffreysi(Weinkauff, 1866) (Haminoeidae), with a discussion on the phylogenetic affinities of the Mediterranean species of the genus. Journal of Natural History, 51: 1593–1608. https://doi.org/10.1080/00222933.2017.1340526

Tibiriça, Y. & Malaquias, M. A. E. 2017. The bubble snails (Gastropoda, Heterobranchia) of Mozambique: an overlooked biodiversity hotspot. Marine Biodiversity, 47(3), 791-811. DOI: 10.1007/s12526-016-0500-7

Malaquias, M. A. E., Zamora-Silva, A., Vitale, D., Spinelli. A., De Matteo, S., Giacobbe, S., Ortigosa, D. & Cervera J. L. 2017. The Suez Canal as a revolving door for marine species: a reply to Galil et al. (2016). Aquatic Invasions, 12(1): 1–4. DOI: https://doi.org/10.3391/ai.2017.12.1.01

Garabedian, K., Malaquias, M.A.E., Crocetta, F., Zenetos, A., Kavadas, S. & Valdés, Á. 2017.Haminoea orteai Talavera, Murillo and Templado, 1987 (Mollusca: Gastropoda: Heterobranchia: Cephalaspidea), a widespread species in the Mediterranean and northeastern Atlantic. Cahiers de Biologie Marine, 58(1): 107-113. DOI: 10.21411/CBM.A.19015AD4

Malaquias, M. A. E., Ohnheiser, L. T., Oskars, T. R. & Willassen, E. 2017. Diversity and systematics of philinid snails (Gastropoda: Cephalaspidea) in West Africa with remarks on the biogeography of the region. Zoological Journal of the Linnean Society, 180(1): 1-35. https://doi.org/10.1111/zoj.12478




Cruz-Rivera, E. & Malaquias, M. A. E. 2016. Ecosystem alterations and species range shifts: An Atlantic-Mediterranean cephalaspidean gastropod in an inland Egyptian lake. PLoS ONE, 11(6): e0156760.

Malaquias, M. A. E, Madrenas, E, & Ballesteros, M. 2016. First occurrence of the tropical Indo-West Pacific head-shield sea slug Chelidonura fulvipunctata in the Balearic Islands confirms its range extension into the western Mediterranean Sea. Mediterranean Marine Science 17 (New Mediterranean Biodiversity Records, March 2016): 232. 

Malaquias, M. A. E., Zamora-Silva, A., Vitale, D., Spinelli. A., De Matteo, S., Giacobbe, S., Ortigosa, D. & Cervera J. L. 2016. The Mediterranean Sea as a gateway for invasion of the Red Sea: The case of the Indo-West Pacific head-shield slug Chelidonura fulvipunctata Baba, 1938. Aquatic Invasions, 11(3): 247–255.

Padula, V., Bahia, J., Stöger, I., Camacho-García, Y., Malaquias, M. A. E., Cervera, J. L., & Schrödl, M. 2016. A test of color-based taxonomy in nudibranchs: molecular phylogeny and species delimitation of the Felimida clenchi (Mollusca: Chromodorididae) species complex.Molecular Phylogenetics and Evolution, 103: 215–229.

Kocot KM, Struck TH, Merkel J, Waits DS, Todt C, Brannock PM, Weese DA, Cannon JT, Moroz LL, Lieb B, Halanych KM. 2016. Phylogenomics of Lophotrochozoa with consideration of systematic error. Systematic Biology, in press.

Passos FD, Corrêa PVF, Todt C. 2016. A new species of Falcidens (Mollusca, Aplacophora, Caudofoveata) from the southeastern Brazilian coast: external anatomy, distribution, and comparison with Falcidens caudatus (Heath, 1918) from the USA. Marine Biodiversity, in press. 

Bergmeier FS, Haszprunar G, Todt C, Jörger KM. 2016. Lost in a taxonomic Bermuda Triangle: comparative 3D-microanatomy of cryptic mesopsammic Solenogastres (Mollusca). Organisms, Diversity & Evolution, in press.

Willassen, E., Williams, A. B., & Oskars, T. R. (2016). New observations of the enigmatic West African Cellana limpet (Mollusca: Gastropoda: Nacellidae).Marine Biodiversity Records, 9(1), 60.

Zamora, B. A. & Malaquias, M. A. E. 2016. Diet preferences of the Aglajidae: a family of cephalaspidean gastropod predators on tropical and temperate shores. Journal of the Marine Biological Association of the United Kingdom, 96(5): 1101–1112.



Caballer, M., Ortea J., Rivero, N., Carias, G., Malaquias, M. A. E. & Narciso, S. 2015. The opisthobranch gastropods (Mollusca: Heterobranchia) from Venezuela: an annotated and illustrated inventory of species. Zootaxa, 4034: 201–256.

Eilertsen, M. E. & Malaquias, M. A. E. 2015. Speciation in the dark: diversification and biogeography of the deep-sea gastropod genus Scaphander in the Atlantic Ocean. Journal of Biogeography, 42: 843–855.

Scherholz M, Redl E, Wollesen T, Todt C, Wanninger A. 2015. From complex to simple: myogenesis in an aplacophoran mollusk reveals key traits in aculiferan evolution. BMC Evolutionary Biology, 15: 201.

Wollesen T, Rodríguez Monje SV, Todt C, Degnan BM, Wanninger A. 2015. Ancestral role of Pax2/5/8 in molluscan
brain and multimodal sensory system development. BMC Evolutionary Biology, 15: 231.

Oskars, T. R., Bouchet, P., & Malaquias, M. A. E. 2015. A new phylogeny of the Cephalaspidea (Gastropoda: Heterobranchia) based on expanded taxon sampling and gene markers. Molecular Phylogenetics and Evolution, 89: 130–150.



Malaquias, M. A. E., Calado, G., Cruz, J. F., and Jensen, K. 2014. Opisthobranch molluscs of the Azores:results of the IV International Workshop on Malacology and Marine Biology (4–13 July 2011) (Mosteiros, Säo Miguel, Azores). Açoreana, Suplemento 10: 139-147.

Malaquias, M.A.E. 2014. New data on the heterobranch gastropods (“opisthobranchs”) for the Bahamas (tropical western Atlantic ocean). Marine Biodiversity Records, 7, e27.

Osca D, Irisarri I, Todt C, Grande C, Zardoya R. 2014. The complete mitochondrial genome of Scutopus ventrolineatus (Mollusca: Chaetodermomorpha) supports the Aculifera hypothesis. BMC Evolutionary Biology, 14: 197.

Redl E, Scherholz M, Todt C, Wollesen T, Wanninger A. 2014. Development of the nervous system in Solenogastres (Mollusca) reveals putative ancestral spiralian features. Evolution and Development, 5: 48. 17 pp.

Kocot KM, Todt C. 2014. Three new meiofaunal solenogaster species (Mollusca: Aplacophora) from the north-east Pacific. Journal of Natural History, 48: 3007-3031.

Portnova D.A., Mokievsky V.O., Haflidason H., Todt C. 2014. Metazoan meiobenthos and nematode assembladges in the Nyegga region of methane seepage (Norwegian Sea). Russian Journal of Marine Biology, 40: 255-265.

Todt C., Kocot KM. 2014. New records on Proneomenia sluiteri (Mollusca: Solenogastres) from Iceland and description of a new species, the brooding Proneomenia custodiens n. sp.. Polish Polar Research, 35: 291-310.

Mikkelsen N.T., Todt C. 2014. Diversity of Caudofoveata (Mollusca) around Iceland and description of Psilodens balduri sp. n.. Polish Polar Research, 35: 279-290.

Ohnheiser, L. T. & Malaquias, M. A. E. 2014. The family Diaphanidae (Gastropoda: Heterobranchia: Cephalaspidea) in Europe, with a redescription of the enigmatic speciesColobocephalus costellatus M. Sars, 1870. Zootaxa, 3774(6): 501-522.

Too, C. C, Carlson, C., Hoff, P. J. and Malaquias, M. A. E. 2014. Diversity and systematics of Haminoeidae gastropods (Heterobranchia: Cephalaspidea) in the tropical West Pacific: new data on the genera AliculastrumAtysDiniatys and LiloaZootaxa, 3794(3): 355-392.



Amorim, A., Oliveira, E., Malaquias, M. A. E. & Lobo-da-Cunha, A. 2013. New insights into the functional morphology of the male copulatory apparatus of bullid gastropods. Zoomorphology, 132: 145-155.

Scherholz M., Redl E., Todt C., Wollesen T., Wanninger A. 2013. Aplacophoran mollusks evolved from ancestors with polyplacophoran-like features. Current Biology, 23:1-5.

Todt C. 2013. Aplacophoran mollusks –Still obscure and difficult? (From the “Mollusks: The Great Unanswered Questions. The James H. Lee Memorial Symposium” presented at 77th Annual Meeting of the American Malacological Society on 24 July 2011 in Pittsburgh, Pennsylvania.) American Malacological Bulletin, 31: 1-7.

Cordeiro, R., Malaquias, M.A.E.,  Mas, G., Figueroa, I., Borges, J. P. & Ávila, S. P. 2013. New records for the opisthobranch fauna of the Archipelago of the Azores (NE Atlantic Ocean). Marine Biodiversity Records, 6:e28.

Eilertsen, M. E. & Malaquias, M. A. E. 2013. Systematic revision of the genusScaphander (Gastropoda, Cephalaspidea) in the Atlantic Ocean with a molecular phylogenetic hypothesis. Zoological Journal of the Linnean Society, 167(3): 389-429.

Eilertsen, M. E. & Malaquias, M. A. E. 2013. Unique digestive system, trophic specialization, and diversification in the deep-sea gastropod genus ScaphanderBiological Journal of the Linnean Society, 109(3): 512-525.

Hanson, D., Cooke, S., Hirano, Y., Malaquias, M. A. E., Crocetta, F., & Valdés, Á. 2013. Slipping through the cracks: The taxonomic impediment conceals the origin and dispersal of Haminoea japonica, an invasive species with impacts to human health. PLoS ONE, 8(10): e77457.

Ohnheiser, L. T. & Malaquias, M. A. E. 2013. Systematic revision of the gastropod family Philinidae (Mollusca: Cephalaspidea) in the northeast Atlantic Ocean with emphasis on the Scandinavian peninsula. Zoological Journal of the Linnean Society, 167(2): 273-326.

Zabbey, N. & Malaquias, M. A. E. 2013. Epifauna diversity and ecology on intertidal flats in the tropical Niger Delta, with remarks on the gastropod species Haminoea orbignyanaJournal of the Marine Biological Association of the United Kingdom, 93(1): 249-257.



Malaquias, M. A. E., Calado, G., Cruz, J. F. & Jensen, K. 2012. On the occurrence of the Caribbean sea slug Thuridilla mazda in the eastern Atlantic Ocean. Marine Biodiversity Records, Vol. 5, e50.

Kingsley R. J., Froelich J., Marks C.B., Spicer L.M., Todt C. 2012. Formation and morphology of epidermal sclerites from a deep-sea hydrothermal vent solenogaster (Helicoradomenia sp., Mollusca). Zoomorphology, 132: 1-9.

Faller S., Rothe B.H., Todt C., Schmidt-Rhaesa A., Loesel R. 2012. Comparative neuroanatomy of the minor molluscan taxa (Caudofoveata, Solenogastres, Polyplacophora, and Scaphopoda) and its phylogenetic implications. 
Zoomorphology, 131: 149-170.



Carmona, L., Malaquias, M. A. E., Gosliner, T. M., Pola, M. & Cervera, J. L. 2011. Amphi-Atlantic distributions and cryptic species in sacoglossan sea slugs. Journal of Molluscan Studies, 77(4): 401-412.

Krylova E.M., Gebruk A.V., Portnova D.A., Todt C., Haflidason H. 2011. New species of the genus Isorropodon (Bivalvia: Vesicomyidae: Pliocardiidae) from cold methane seeps at Nyegga (Norwegian Sea, Vøring Plateau, Storrega slide). Journal of the Marine Biological Association of the United Kingdom, 91 (5): 1135-1144.

Malaquias, M. A. E., Gosliner, T. M. & Cervera, J. L. 2011. Studies on opisthobranch molluscs: Proceedings of a symposium, World Congress of Malacology, Phuket, Thailand, July 2010. Introduction. Journal of Molluscan Studies, 77(4): 321.

Pedro, N. C., Malaquias, M. A. E., Costa, A. C. & Ávila, S. P. 2011. Crimora papillata (Nudibranchia: Triophinae), a new record for the shallow marine molluscs of the Azores. Marine Biodiversity Records, 4: e37.

Zenetos, A., Katsanevakis, S., Poursanidis, D., Crocetta, F., Damalas, D., Apostolopoulos, G., Gravili, C., Vardala-Theodorou, E. & Malaquias, M. 2011. Marine alien species in Greek Seas: Additions and amendments by 2010.Mediterranean Marine Science, 12(1): 95-120.



Meyer, A., Todt, C., Mikkelsen, N. T., Lieb, B. 2010. Fast evolving 18S rRNA sequences from Solenogastres (Mollusca) resist standard PCR amplification and give new insights into mollusk substitution rate heterogeneity. BMC Evolutionary Biology 10.

Schander, C., Rapp, H. T., Kongsrud, J. A., Bakken, T., Berge, J., Cochrane, S., Oug, E., Byrkjedal, I., Todt, C., Cedhagen, T. Fosshagen, A., Gebruk, A. V., Larsen, K., Levin, L., Obst, M., Pleijel, F., Stöhr, S., Warén, A., Mikkelsen, N. T., Hadler-Jacobsen, S., Keuning, R., Petersen, K. H., Thorseth, I. H., Pedersen, R. 2010. The fauna of hydrothermal vents on the Mohn Ridge (North Atlantic). Marine Biology Research6. 155–171.

Malaquias, M. A. E. 2010. Systematics, phylogeny, and natural history of Bullacta exarata (Philippi, 1849): an endemic cephalaspidean gastropod from the China Sea. Journal of Natural History, 44(33/34): 2015–2019.



Ivanov, D. L., Mikkelsen, N. T., Schander, C. 2009. Falcidens sagittiferusSalvini-Plawen, 1968: additional data on morphology and distribution (Mollusca, Aplacophora, Caudofoveata). Fauna Norvegica, 29: 3–9.

Ciavatta, M. L., Manzo, E., Nuzzo, G. Villani, G., Cimino, G., Cervera, J. L., Malaquias, M. A. E. & Gavagnin, M. 2009. Aplysiopsenes: an additional example of marine polyketides with a mixed acetate/propionate pathway. Tetrahedron Letters, 50: 527-529.

Malaquias, M. A. E. & Reid, D. G. 2009. Tethyan vicariance, reliction and speciation: evidence from a global molecular phylogeny of the opisthobranch genus BullaJournal of Biogeography, 36: 1760-1777.

Malaquias, M. A. E., Bericibar, E., & Reid, D. G. 2009. Reassessment of the trophic position of Bullidae (Gastropoda: Cephalaspidea) and the importance of diet in the evolution of cephalaspidean gastropods. Journal of Zoology, 277: 88-97.

Malaquias, M. A. E., Dodds, J. M., Bouchet, P., & Reid, D. G. 2009. A molecular phylogeny of the Cephalaspidea sensu lato (Gastropoda: Euthyneura): Architectibranchia redefined and Runcinacea reinstated. Zoologica Scripta, 38(1): 23-41.

Malaquias, M. A. E., Calado, G. P., Padula, V., Villani, G. & Cervera, J. L. 2009. Molluscan diversity in the North Atlantic Ocean: New records of opisthobranchs gastropods from the Archipelago of the Azores. Marine Biodiversity Records, 2: e38.



Malaquias, M. A. E & Reid, D. G. 2008. Functional morphology of the gonoduct of the gastropod Bulla striata Bruguière, 1792 (Opisthobranchia: Cephalaspidea): evidence for a monaulic system. Acta Zoologica, 89: 205-210.

Malaquias, M. A. E. & Reid, D. G. 2008. Systematic revision of the Recent species of Bullidae (Mollusca: Gastropoda: Cephalaspidea), with a molecular phylogenetic analysis. Zoological Journal of the Linnean Society, 153: 453–543.