Centre for Cancer Biomarkers CCBIO
CCBIO Research School

BMED904 - Graduate Course in Extracellular Matrix

The course will cover various aspects of extracellular matrix (ECM) biology. A recurring theme will be the roles of the various ECM molecules and their functions in health and disease. Note, due to the uncertainty regarding the Covid-19 situation, the course is planned to combine face-to-face “seat time” for students attending in Bergen with online presentations of all lectures and lab demonstrations. The lectures are open to all interested.

Microscopic photo.
P. M. Erusappan

The PhD course Matrix Biology (BMED904) course will run June 07-11, 2021.

Place: option of on location at the University of Bergen, Haukeland University Hospital, or through a live Zoom streaming online.

This year, the BMED904 course will be joint with the DIKU Summer School in Fibrosis.


  • Extracellular matrix
  • Imaging in biomedical research
  • Proteoglycans, cell-matrix receptors
  • Laminin- muscular dystrophy
  • The brain matrix and its potential role in disease
  • Fibroblast heterogeneity
  • Artificial matrices for stem cell culture 
  • Fibroblasts in fibrosis research
  • Fibroblast cell migration
  • Traction Force microscopy
  • Myofibroblasts
  • Collagens and cell-collagen interactions
  • Methods for discovery of new proteins that mediate remodeling of the extracellular matrix
  • Heart Fibrosis 
  • The multifaceted role of CCN proteins in fibrotic stroma
  • Integrins in fibrosis
  • Mechanotransduction in cancer
  • CAF heterogeneity 


  • Wayne CarverUSC, USA 
  • Daniela E. Costea, UiB, Norway
  • Martin Decaris, Pliant Therapeutics, USA
  • Madeleine Durbeej- Hjalt, Lund univ. Sweden
  • Donald Gullberg, UiB, Norway
  • Ritva Heljasvaara, Univ. Oulu, Finland
  • Marion Kusche-Gullberg, UiB, Norway
  • Andrew Leask, USask, Canada
  • Chris McCulloch, UoT, Canada
  • Cathy Merry, Nottingham UK
  • Roya Navab, UoT, Canada
  • Roger Oria-Fernandez, UCSF, USA 
  • Joanna Phillips, UCSF, USA               
  • Sergey Plotnikov, UoT, Canada
  • Andreas Romaine, UiO, Norway
  • Ronen Schuster, UoT, Canada
  • Kalle Sipilä, London, UK
  • Frits ThorsenUiB, Norway

Scientific program: See the preliminary program at this link.

ECTS: This is a 3 ECTS course at the University of Bergen. NOTE: If you need to register ECTS, you need to either be a registered student at the UiB, or register as a guest student.

Registration: Students at the UiB register through Studentweb. Deadline is February 1st. External students who would like to have the ECTS added, register through Søknadsweb, where they simultaneously apply for UiB guest student status. Deadline is February 1st. Participants only interested in joining the Zoom lectures for professional updates and don’t need the ECTS, register through another form, soon to be added here. 

Contact: Administrative coordinator is Margarethe Bittins. Academic responsible are Marion Kusche-Gullberg and Donald Gullberg.